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Fir

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From Wikipedia, the free encyclopedia
Genus of coniferous trees

For other uses, seeFIR (disambiguation) andFIRS (disambiguation).
"Fir tree" redirects here. For other uses, seeFir Tree (disambiguation).

Fir
Cones of a fir tree
Korean fir (Abies koreana) cones and foliage
Scientific classificationEdit this classification
Kingdom:Plantae
Clade:Tracheophytes
Clade:Gymnospermae
Division:Pinophyta
Class:Pinopsida
Order:Pinales
Family:Pinaceae
Subfamily:Abietoideae
Genus:Abies
Mill.
Type species
Abies alba
Species

See text

Synonyms
  • PeuceRichard 1810

Firs areevergreenconiferous trees belonging to thegenusAbies (Latin:[ˈabieːs]) in the familyPinaceae. There are approximately 48–65[2][3] extant species, found on mountains throughout much of North andCentral America,Eurasia, andNorth Africa. The genus is most closely related toKeteleeria, a small genus confined to eastern Asia.[4]

They are tall trees that can be distinguished from other members of the pine family by the way in which their needle-likeleaves are attached singly to the branches with a circular base, and by theircones, which, like those ofcedars, stand upright on the branches like candles and disintegrate at maturity.

The wood of firs is used forpulp to make paper, forplywood, and for indoor construction. Some species serve asChristmas trees, while others are used as decorative trees with their brightly coloured cones. In art,Lucas Cranach the Elder paintedMadonna under the fir tree forWrocław Cathedral in 1510.

Etymology

[edit]

The English name "fir" derives from the Old Norsefyri or the Old Danishfyr.[5] The generic nameAbies is the Latin for "fir".[6]

Description

[edit]

Fir trees are tall, often 40–60 metres and sometimes approaching 100 metres high, usually with a single straight trunk. The crown starts conical, but becomes more varied in shape with age. The primary branches are arranged in whorls around the trunk.[7]

Leaves

[edit]

Firs have needle-likeleaves, arranged spirally but often appearing to be in two or more rows on opposite sides of the twigs. The base of each leaf is round and attached to a small pit in the twig. Each leaf is normally twisted at its base so that the side withstomata faces downwards.[7] In the upper crown on cone-bearing branches, the leaves are shorter, curved, and sometimes sharp.[8]

  • A. alba – the underside of leaves have two whitish strips formed by wax-covered stomatal bands, while their bases are shaped like suction cups.
    A. alba – the underside of leaves have two whitish strips formed bywax-coveredstomatal bands, while their bases are shaped like suction cups.
  • A. grandis foliage – upper side of the leaves, showing the leaves lying flat either side of the shoot
    A. grandis foliage – upper side of the leaves, showing the leaves lying flat either side of the shoot
  • Foliage of A. pinsapo showing the radial leaf arrangement in this species
    Foliage ofA. pinsapo showing the radial leaf arrangement in this species
  • Most firs are in between flat and radial, often with longer leaves at the sides, and shorter leaves above the shoot; here, A. mariesii in Japan
    Most firs are in between flat and radial, often with longer leaves at the sides, and shorter leaves above the shoot; here,A. mariesii in Japan
  • Cross-section of needle leaf of Abies nordmanniana
    Cross-section of needle leaf ofAbies nordmanniana

Cones

[edit]

Firs differ from other conifers such asspruces in having erect, cylindrical female seedcones 5–25 cm (2–10 in) long that disintegrate at maturity to release the wingedseeds. Seed cones take a year to become mature; they start out green or other bright colour, darkening as they develop to dark brown or black. The leaflike seedbracts are visible when young, and in some species remain so. The seeds sit in thin cups; each seed has a triangle-shaped wing. The male cones are pendulous (dangling) and resemblecatkins; both pollen and seeds are wind-dispersed.[7]

  • Intact and disintegrated Bulgarian fir cones
    Intact and disintegratedBulgarian fir cones
  • Immature cones of some species are green; here Manchurian fir Abies holophylla
    Immature cones of some species are green; hereManchurian firAbies holophylla
  • Disintegrating cones of Manchurian fir
    Disintegrating cones of Manchurian fir
  • Pindrow fir Abies pindrow with dark purple cones
    Pindrow firAbies pindrow with dark purple cones
  • Caucasian Fir Abies nordmanniana young cones with reddish scales and yellow-green bracts
    Caucasian FirAbies nordmanniana young cones with reddish scales and yellow-green bracts
  • Noble fir Abies procera, with five heavy (20 cm, approx 0.5 kg each) seed cones
    Noble firAbies procera, with five heavy (20 cm, approx 0.5 kg each) seed cones
  • Pollen cones
    Pollen cones

Evolution

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Fossil history

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FossilAbies milleri foliage and axis.Early Eocene, 49.5 mya

The oldest pollen assignable to the genus dates to theLate Cretaceous inSiberia, with records of leaves and reproductive organs across the Northern Hemisphere from theEocene onwards.[9]

External phylogeny

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Based ontranscriptome analysis,Keteleeria is sister toAbies, with the Pseudolariceae the next nearest relatives.[10]

Pinaceae
Abietoideae
Cedreae

Cedrus

Pseudolariceae
Abieteae
(firs and allies)
Pinoideae

(pines, larches, etc)

Internal phylogeny

[edit]

Phylogeny ofAbies based on phylogenomic analysis in 2021:[11][12]

Abies
(Pseudotorreya)

A. bracteata(Don) Poit.

(Religio)
section

A. mariesiiMasters

A. amabilis(Douglas ex Loudon) Forbes

A. proceraRehder

Amabilis
section

A. magnificaMurray

Nobiles
section

A. concolor(Gordon) Lindley ex Hildebr.

A. jaliscana(Martínez) Mantilla, Shalisko & Vázquez

A. guatemalensisRehder

A. hickeliiFlous & Gaussen

A. flinckiiRushforth

A. vejariiMartínez

A. durangensisMartínez

A. religiosa(Kunth) von Schlechtendal & von Chamisso

A. hidalgensisDebreczy, Rácz & Guízar

A. grandis(Douglas ex Don) Lindley

A. lowiana(Gordon) Murray

Grandis
(Abies)
section

A. albaMiller

A. pinsapoBoiss.

A. cephalonicaLoudon

A. nebrodensis(Lojac.) Mattei

A. nordmanniana(Steven) Spach

A. numidicade Lannoy ex Carrière

A. ×borisii-regisMattf.

A. cilicica(Antoine & Kotschy) Carrière

Abies
section

A. lasiocarpa(Hooker) Nuttall

A. ernestiiRehder

A. balsamea(von Linné) Miller

Balsamea
section

A. firmaSiebold & Zuccarini

A. sibiricaLedeb.

Sibiria
section

A. fanjingshanensisHuang, Tu & Fang

A. ziyuanensisFu & Mo

series

A. kawakamii(Hayata) Itô

A. chensiensisvan Tieghem

A. squamataMasters

A. beshanzuensisWu

A. pindrow(Royle ex Don) Royle

Squamatae
series

A. koreanaWilson

A. nephrolepis(Trautvetter ex Maxim.) Maxim.

A. holophyllaMaxim.

A. sachalinensis(Schmidt) Masters

A. fabri(Masters) Craib

A. veitchiiLindley

A. fraseri(Pursh) Poiret

A. delavayiFranchet

A. densaGriffith

A. spectabilis(Don) de Mirbel

A. nukiangensisCheng & Fu

A. ferreanaBordères & Gaussen

A. forrestiiColtm.-Rog.

A. georgeiOrr

A. homolepisSiebold & Zuccarini

A. yuanbaoshanensisLu & Fu

Spectabiles
Pseudopicea

Taxonomy

[edit]

Taxonomic history

[edit]

In 1754,Philip Miller set up the genusAbies; he also defined the type speciesAbies alba.[13][14] In 1878,George Engelmann classified only a part of the genus;[13]Heinrich Mayr did the same in 1890,[13][15] as did the German botanist Wilhelm Patschke in 1913.[13] The classifications by Paul Robert Hickel in 1906 to 1908,[13][16] and by P. Landry in 1984,[13][17] made use only of a subset of the available morphological characteristics.[13] In 1990 and 2001, the Dutch botanistAljos Farjon attempted a more complete classification; he accepted 48 species within the genus; in 2017 he reduced this to 46. Adopting a different approach, in 2011 Z. Debreczy and I. Rácz treated the genus as containing 67 species.[7]

Species

[edit]

As of September 2025[update],Plants of the World Online accepted 49 species.[18] The sections are based on Stull et al. 2021.[11]

SectionAbies is found in central, south, and eastern Europe and Asia Minor.

SectionBalsamea is found in northern Asia and North America, and high mountains further south.

SectionGrandis is found in western North America to Mexico, Guatemala, Honduras and El Salvador, in lowlands in the north, moderate altitudes in south.

  • Abies grandis – grand fir or giant fir
    • Abies grandis var.grandis – Coast grand fir
    • Abies grandis var.idahoensis – interior grand fir
  • Abies concolor – white fir
    • Abies concolor subsp.concolor – Rocky Mountain white fir or Colorado white fir
    • Abies concolor subsp.lowiana – Low's white fir or Sierra Nevada white fir
  • Abies durangensis – Durango fir
    • Abies durangensis var.coahuilensis – Coahuila fir
  • Abies flinckii – Jalisco fir
  • Abies guatemalensis – Guatemalan fir
    • Abies guatemalensis var.guatemalensis
    • Abies guatemalensis var.jaliscana
  • Abies vejarii

SectionMomi is found in east and central Asia and the Himalaya, generally at low to moderate altitudes.

SectionAmabilis is found in the Pacific Coast mountains in North America and Japan, in high rainfall areas.

A. fabri, Sichuan, China

SectionPseudopicea is found in theSino – Himalayan mountains at high altitudes.

SectionOiamel is found in central Mexico at high altitudes.

A. magnifica, California

SectionNobilis (westernU.S., high altitudes)

SectionBracteata (California coast)

SectionIncertae sedis

Ecology

[edit]

Distribution and habitat

[edit]

Firs are distributed around the Northern Hemisphere. The genus is native across much of North America, Eurasia, Turkey, Syria, Morocco, and Algeria. It is introduced in Scandinavia and the British Isles.[21]Abies sibirica grows as far north as 67°N in Siberia, whileA. guatemalensis grows as far south as 15°N in Central America. Most firs favour cold climates, whether at altitude in mountain ranges or at high latitude. Many species haverelict distributions, occupying small areas of what were once much larger distributions. Only a few species are widespread.[7]

Abies religiosa (sacred fir) trees giveroosting shelter to overwinteringmonarch butterflies.[22][23]

Pests and diseases

[edit]

Firs are hosts to a variety of invertebrate pests and fungal diseases. Pest groups includeadelgid bugs,aphids,bark beetles,clearwing moths,conifer twig weevils,caterpillars of some moths,nematodes,sawflies,spider mites, andspittlebugs. Diseases of firs include annosus (Heterobasidion)root rot,cankers, andneedle cast.[24]

Uses

[edit]

Unlike theDouglas fir (Pseudotsuga), firs producesoftwood, often used aspulp or for the manufacture ofplywood and rough timber. It is commonly used inCanadian Lumber Standard graded wood, used for internalstud walls and similar applications.[25]Abies spectabilis is used inAyurveda as anantitussive (cough suppressant) drug.[26] Firs produce a variety ofterpenoids that could have practical uses. Terpenoid composition of the bark varies by genetics, geography, age and size of the tree.[27][28]

Caucasian fir,noble fir,Fraser's fir andbalsam fir are popularChristmas trees, generally considered to be the best for this purpose, with aromatic foliage that does not shed many needles on drying out. Many form decorative garden trees, notablyKorean fir and Fraser's fir, which produce brightly coloured cones even when very young, still only 1–2 m (3 ft 3 in – 6 ft 7 in) tall. Many fir species are grown in botanic gardens and other specialist tree collections in Europe and North America.[29]

In art

[edit]

Lucas Cranach the Elder paintedMadonna under the fir tree forWrocław Cathedral in 1510.[30] The painting was taken to Germany after the Second World War for restoration, but a copy was made and substituted for the original. The forgery was discovered, partly because the copy was on a fir board where the original was onlime wood. The original was returned to Poland in 2012, the most valuable painting to be restituted to the country.[31]

See also

[edit]

References

[edit]
  1. ^abSchorn, Howard; Wehr, Wesley (1986)."Abies milleri, sp. nov., from the Middle Eocene Klondike Mountain Formation, Republic, Ferry County, Washington".Burke Museum Contributions in Anthropology and Natural History.1:1–7.
  2. ^Torres, Leon Nahuel; Shi, Xiao; Na, Yuling; Wang, Bing; Tian, Chi; Chen, Jun (2024-03-01)."First study on fossil wood from the Middle Pleistocene of the Songliao Plain, Northeast China".Review of Palaeobotany and Palynology.322 105063.Bibcode:2024RPaPa.32205063T.doi:10.1016/j.revpalbo.2024.105063.ISSN 0034-6667.
  3. ^Debreczy, Zsolt; Rácz, István; Musial, Kathy (2011).Conifers Around the World : Conifers of the Temperate Zones and Adjacent Regions. Budapest: DendroPress.ISBN 978-9632190617.
  4. ^Leslie, Andrew B.; et al. (2018)."ajb21143-sup-0004-AppendixS4"(PDF).American Journal of Botany.105 (9):1531–1544.doi:10.1002/ajb2.1143.PMID 30157290.
  5. ^"fir".Online Etymology Dictionary.Archived from the original on 2021-08-08. Retrieved2020-10-01.
  6. ^"Abies, subst. masc".Centre national de ressources textuelles et lexicales. Retrieved8 September 2025.
  7. ^abcde"Abies Mill". International Dendrology Society. Retrieved8 September 2025.
  8. ^Arno, Stephen F.; Hammerly, Ramona P. (2020) [1977].Northwest Trees: Identifying & Understanding the Region's Native Trees (field guide ed.). Seattle:Mountaineers Books. p. 125.ISBN 978-1-68051-329-5.OCLC 1141235469.Archived from the original on 2023-09-19. Retrieved2022-02-26.
  9. ^Xiang, Xiaoguo; Cao, Ming; Zhou, Zhekun (October 2007)."Fossil history and modern distribution of the genus Abies (Pinaceae)".Frontiers of Forestry in China.2 (4):355–365.doi:10.1007/s11461-007-0058-4.Archived from the original on 2024-01-01. Retrieved2023-02-15.
  10. ^Stull, Gregory W.; Qu, Xiao-Jian; Parins-Fukuchi, Caroline; Yang, Ying-Ying; Yang, Jun-Bo; et al. (19 July 2021)."Gene duplications and phylogenomic conflict underlie major pulses of phenotypic evolution in gymnosperms".Nature Plants.7 (8):1015–1025.Bibcode:2021NatPl...7.1015S.doi:10.1038/s41477-021-00964-4.PMID 34282286.S2CID 236141481.
  11. ^abStull, Gregory W.; Qu, Xiao-Jian; Parins-Fukuchi, Caroline; Yang, Ying-Ying; Yang, Jun-Bo; et al. (2021)."Gene duplications and phylogenomic conflict underlie major pulses of phenotypic evolution in gymnosperms".Nature Plants.7 (8):1015–1025.Bibcode:2021NatPl...7.1015S.bioRxiv 10.1101/2021.03.13.435279.doi:10.1038/s41477-021-00964-4.PMID 34282286.S2CID 232282918.
  12. ^Stull, Gregory W.; et al. (2021)."main.dated.supermatrix.tree.T9.tre". Figshare.doi:10.6084/m9.figshare.14547354.v1.Archived from the original on 2024-01-01. Retrieved2023-03-30.
  13. ^abcdefgFarjon, Aljos; Rushworth, K. D. (1989)."A Classification of Abies Miller (Pinaceae)"(PDF).Notes of the Royal Botanic Garden Edinburgh.46 (1): 59.doi:10.24823/nrbge.1989.3565.
  14. ^Miller, Philip (1754).The Gardener's Dictionary. Vol. 1 (4th, abridged ed.).
  15. ^Mayr, Heinrich (1890).Monographie der Abielineen des Japanischen Reiches. Munchen.{{cite book}}: CS1 maint: location missing publisher (link)
  16. ^Hickel, Robert (1906-08). Notes pour servir ä la determination pratique des Abietinees. Bull. Soc. Dendrol. France 2: 45 58, f. I 7(15 xi 1906); 3; 5 18, f. 8 30 (15 ii 1907); 4: 41 48, f. 31 46(15v 1907); 5: 82 86(15 viii 1907); 7:5-10, f. a 1 (15 ii 1908); 9; 179-185 (15 viii 1908); 10: 201 208 (15 xi 1908).
  17. ^Landry, Pierre (1984)."Synopsis du genre Abies".Bulletin de la Société Botanique de France. Lettres Botaniques (in French).131 (3):223–229.Bibcode:1984BSBFL.131..223L.doi:10.1080/01811797.1984.10824633.
  18. ^"Abies Mill. Accepted Species".Royal Botanic Gardens, Kew. Retrieved10 September 2025.
  19. ^Knowlton, F.H. (1923). Fossil plants from the Tertiary lake beds of South-Сentral Colorado (Report). Professional Paper. Vol. 131-G. United States Geological Survey. pp. 183–197.doi:10.3133/pp131G.
  20. ^Wolfe, J.A.; Schorn, H.E. (1990). Taxonomic revision of the Spermatopsida of the Oligocene Creede flora, southern Colorado (Report). Bulletin. Vol. 1923. United States Geological Survey. pp. 1–40.doi:10.3133/b1923.
  21. ^"Abies Mill. Distribution".Royal Botanic Gardens, Kew. Retrieved10 September 2025.
  22. ^Groth, Jacob (10 November 2000)."Monarch Migration Study". Swallowtail Farms.Archived from the original on 9 May 2022. Retrieved21 July 2014.
  23. ^"Monarch Migration". Monarch Joint Venture. 2013.Archived from the original on 2017-10-31. Retrieved2017-10-25.
  24. ^"Pests in Gardens and Landscapes". University of California Agriculture & Natural Resources: Statewide Integrated Pest Management Program. Retrieved10 September 2025.
  25. ^Jenkins, Steve (2023-09-03)."What is CLS timber and what DIY projects is it good for?".Homebuilding & Renovating. Retrieved2024-08-22.
  26. ^Kershaw, Linda (2000).Edible and Medicinal Plants of the Rockies. Edmonton, Alberta:Lone Pine Publishing. p. 26.ISBN 978-1-55105-229-8.
  27. ^Hemingway, R. W. (2018-01-18). "Bark: Its Chemistry and Prospects for Chemical Utilization".Organic Chemicals from Biomass. CRC Press. pp. 189–248.ISBN 978-1-351-07525-1.
  28. ^Barton, George M. (2018-01-18). "Foliage".Organic Chemicals from Biomass. CRC Press. pp. 249–280.ISBN 978-1-351-07525-1.
  29. ^Christian, T. (2021)."Abies".Trees and Shrubs Online.Archived from the original on 2023-03-01. Retrieved2023-03-01.
  30. ^""Madonna under the Fir Tree" by Lucas Cranach the Elder".National Museum in Wrocław. 22 September 2020. Retrieved9 September 2025.
  31. ^"Madonna under the Fir Tree".The Marian Temples Trail. Retrieved9 September 2025.

Further reading

[edit]

External links

[edit]
Wikimedia Commons has media related toAbies.
Classification ofArchaeplastida orPlantaes.l.
incertae sedis
Glaucoplantae
Glaucophyta
Rhodoplantae
Picozoa
Rhodelphidia
Rhodophyta
(red algae)
Cyanidiophytina
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(green algae & land plants)
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Prasinophytina
Chlorophytina
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(seed ferns)
and other extinct
seed plant groups
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(living gymnosperms)
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(flowering plants)
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