Family of beetles
Endomychidae , orhandsome fungus beetles , is a family ofbeetles with representatives found in allbiogeographic realms . There are around 120 genera and 1300 species. The family was established based on the type genusEndomychus Panzer which was applied to a species that Linnaeus calledChrysomela coccinea . As the common name suggests, Endomychidae feed onfungi .Crowson , in his influential treatment of the beetles, placed the family within theCucujoidea . They have a tarsal formal of 4-4-4 or 3-3-3 and the wings lack a closed radial cell. The second antennal segment has a sensory appendage that is as long as the third antennal segment. The family has also been grouped with theCoccinellidae in a group called the Trimera for having pseudotrimerous tarsi.[ 1] Anamorphidae . Mycetaeinae and Eupsilobiinae were also found not to belong within theclades of the core Endomychidae, and likewise reclassified into the familiesMycetaeidae andEupsilobiidae .[ 2]
Thesubfamilies that are included:
These 70 genera belong to the family Endomychidae as defined in the past (due to revisions, this may not be entirely accurate):
Acinaces Gerstaecker, 1858 g Aclemmysa Reitter, 1904 g Agaricophilus Motschulsky, 1838 g Amphisternus Germar, 1843 g Amphistethus Strohecker, 1964 g Ancylopus Costa, 1854 g Anidrytus Gerstaecker, 1858 i c g Aphorista Gorham, 1873 i c g b Archipines Strohecker, 1953 i c g Asymbius Gorham, 1896 g Atopomychus Tomaszewska & Szawaryn, 2013 g Atrichonota Arrow, 1925 g Avencymon Strohecker, 1971 g Beccariola Arrow, 1943 g Bolbomorphus Gorham, 1888 g Brachytrycherus Arrow, 1920 g Bystodes Strohecker, 1953 g Cholovocera Motschulsky, 1838 g Chondria Gorham, 1888 g Corynomalus Chevrolat in Dejean, 1836 i c g Cyclotoma Mulsant, 1851 i g Danae Reiche, 1847 i c g b Danascelis Tomaszewska, 1999 i c g Dapsa Latreille, 1829 g Dexialia Sasaji, 1970 g Dialexia Gorham, 1891 g Discolomopsis Shockley, 2006 g Displotera Reitter, 1877 g Ectomychus Gorham, 1888 g Eidorus g Endomychus Panzer, 1795 i c g b Ephebus Chevrolat in Dejean, 1836 i c g Epipocus Germar, 1843 i c g b Epopterus Chevrolat in Dejean, 1836 i c g Eucteanus Gerstaecker, 1857 g Eucymon g Eumorphus Weber, 1801 g Geoendomychus Lea, 1922 g Glesirhanis Shockley & Alekseev, 2014 g Hadromychus Bousquet & Leschen, 2002 i c g b Holoparamecus Curtis, 1833 i c g b Hylaia Guerin, 1857 g Hylaperdina Tomaszewska, 2012 g Idiophyes Blackburn, 1896 g Indalmus Gerstaecker, 1858 g Leiestes Chevrolat, 1836 g Lycoperdina Latreille, 1807 i c g b Meilichius Gerstaecker, 1857 g Merophysia Lucas, 1852 g Microxenus Wallaston, 1861 g Mycetina Mulsant, 1846 i c g b Mychothenus Strohecker, 1953 g Natalinus Tomaszewska, 2011 g Ohtaius Chûjô, 1938 g Palaeoestes Kirejtshuk & Nel, 2009 g Paniegena Heller, 1916 g Parasymbius Arrow, 1920 g Phymaphora Newman, 1838 i c g b Phymaphoroides Motschoulsky, 1857 g Pleganophorus Hampe, 1855 g Polymus Mulsant, 1846 g Pseudindalmus Arrow, 1920 g Rhanidea Strohecker, 1953 i c g b Saula Gerstaecker, 1858 i c g Sinocymbachus Strohecker & Chujo, 1970 g Stenotarsus Perty, 1832 i c g b Stethorhanis Blaisdell, 1931 i c g Trochoideus Westwood, 1833 i c g b Trycherus Gerstaecker, 1857 g Xenomycetes Horn, 1880 i c g b Data sources: i = ITIS,[ 3] [ 4] [ 5] [ 6]
Burmalestes [ 7] Burmese amber , Myanmar, Late Cretaceous (Cenomanian )Cretolestes [ 7] Cretaparamecus [ 7] Palaeomycetes [ 7] Zemyna [ 7] Laima Alekseev and Tomaszewska, 2018) :Baltic amber , Eocene^ Tomaszewska, K.Wioletta (2000)."Morphology, phylogeny and classification of adult endomychidae (Coleoptera: Cucujoidea)" (PDF) .Annales Zoologici .50 (4):449– 558. ^ Robertson, James A; Ślipiński, Adam; Moulton, Matthew; Shockley, Floyd W; Giorgi, Adriano; Lord, Nathan P; McKenna, Duane D; Tomaszewska, Wioletta; Forrester, Juanita; Miller, Kelly B; Whiting, Michael F; McHugh, Joseph V (2015)."Phylogeny and classification of Cucujoidea and the recognition of a new superfamily Coccinelloidea (Coleoptera: Cucujiformia)" (PDF) .Systematic Entomology .40 (4): 745.Bibcode :2015SysEn..40..745R .doi :10.1111/syen.12138 .S2CID 55206626 . ^ "Endomychidae Report" .Integrated Taxonomic Information System . Retrieved2018-05-07 .^ "Browse Endomychidae" .Catalogue of Life . Archived fromthe original on 2021-06-18. Retrieved2018-05-07 .^ "Endomychidae" .GBIF . Retrieved2018-05-07 .^ "Endomychidae Family Information" .BugGuide.net . Retrieved2018-05-07 .^a b c d e Tomaszewska, Wioletta; Ślipiński, Adam; Bai, Ming; Zhang, Weiwei; Ren, Dong (November 2018)."The oldest representatives of Endomychidae (Coleoptera: Coccinelloidea) from the Upper Cretaceous Burmese amber" Cretaceous Research .91 :287– 298.Bibcode :2018CrRes..91..287T .doi :10.1016/j.cretres.2018.07.001 .S2CID 135401188 .
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