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Didymictinae

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Extinct subfamily of carnivores

Didymictinae Temporal range:63.8–39.7 Ma PreꞒ Ꞓ O S D C P T J K Pg N earlyPaleocene - middleEocene

skull ofDidymictis protenus
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Clade:	Carnivoramorpha
Superfamily:	†Viverravoidea
Family:	†Viverravidae
Subfamily:	†Didymictinae Flynn & Galiano, 1982^[1]
Type genus
†Didymictis Cope, 1875
Genera
†Bryanictis †Didymictis †Intyrictis †Pristinictis †Protictis^{(paraphyletic genus)} †Raphictis Incertae sedis: †"Deltatherium" durini
Synonyms
Didymictida(Flynn & Galiano, 1982) Didymictidae(Flynn & Galiano, 1982)

Didymictinae ("doubleweasels") is anextinct subfamily of mammals from extinct familyViverravidae, that lived from the earlyPalaeocene to the middleEocene inNorth America andEurope.^[2]

Classification and phylogeny

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Classification

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Subfamily: †Didymictinae(Wortman & Matthew, 1899)
- Genus: †Bryanictis(MacIntyre, 1966)
  - †Bryanictis microlestes(Simpson, 1935)
  - †Bryanictis paulus(Meehan & Wilson, 2002)
  - †Bryanictis terlinguae(Standhardt, 1986)
- Genus: †Didymictis(Cope, 1875)
  - †Didymictis altidens(Cope, 1880)
  - †Didymictis dellensis(Dorr, 1952)
  - †Didymictis leptomylus(Cope, 1880)
  - †Didymictis protenus(Cope, 1874)
  - †Didymictis proteus(Polly, 1997)
  - †Didymictis vancleveae(Robinson, 1966)
  - †Didymictis sp. [Erquelinnes, Hainaut, Belgium](Dollo, 1909)
- Genus: †Intyrictis(Gingerich & Winkler, 1985)
  - †Intyrictis vanvaleni(MacIntyre, 1966)
- Genus: †Pristinictis(Fox & Youzwyshyn, 1994)
  - †Pristinictis connata(Fox & Youzwyshyn, 1994)
- Genus: †Protictis(Matthew, 1937)
  - †Protictis agastor(Gingerich & Winkler, 1985)
  - †Protictis haydenianus(Cope, 1882)
  - †Protictis minor(Meehan & Wilson, 2002)
  - †Protictis paralus(Holtzman, 1978)
  - †Protictis simpsoni(Meehan & Wilson, 2002)
  - †Protictis aprophatos(Flynn & Galiano, 1982)
- Genus: †Raphictis(Gingerich & Winkler, 1985)
  - †Raphictis gausion(Gingerich & Winkler, 1985)
  - †Raphictis iota(Scott, 2008)
  - †Raphictis machaera(Rankin, 2009)
  - †Raphictis nanoptexis(Rankin, 2009)
- Incertae sedis:
  - †"Deltatherium" durini(Van Valen, 1978)

References

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^J. J. Flynn and H. Galiano. (1982.) "Phylogeny of Early Tertiary Carnivora, With a Description of a New Species ofProtictis From the Middle Eocene of Northwestern Wyoming" American Museum Novitates 2725:1-64
^McKenna, Malcolm C.; Bell, Susan K. (1997).Classification of Mammals Above the Species Level. New York: Columbia University Press.ISBN 978-0-231-11012-9. Retrieved16 March 2015.

Mammals of cladePan-Carnivora

Kingdom:Animalia
Phylum:Chordata
Class:Mammalia
Infraclass:Placentalia
Superorder:Laurasiatheria
Clade:Scrotifera
Grandorder:Ferungulata
Mirorder:Ferae

Pan-Carnivora

Clade:
Pan-Carnivora

Clade:
Carnivoramorpha

SeeCarnivoramorpha below ↓

Order:
†Hyaenodonta

SeeHyaenodonta below ↓

Order:
†Oxyaenodonta

Family:
†Oxyaenidae

Subfamily:
†Machaeroidinae

†Apataelurus †Diegoaelurus †Isphanatherium †Machaeroides
Incertae sedis	†Machaeroidinae sp. (CM 2386) †Machaeroidinae sp. (FMNH PM 1506) †Machaeroidinae sp. (USNM 173514)

Subfamily:
†Oxyaeninae

Subfamily:
†Palaeonictinae

†Ambloctonus
†Dipsalodon^{(paraphyletic genus)}
†Palaeonictis

Subfamily:
†Tytthaeninae

†Tytthaena

other
representatives

†Altacreodus
Family: †Simidectidae	†Simidectes
Family: †Wyolestidae	†Wyolestes

Carnivoramorpha

Clade:
Carnivoramorpha

Clade:
Carnivoraformes
(i.e. Clade "A")

Clade "B"

Family:
†Quercygalidae

†Quercygale

†Gracilocyon/Oodectes
clade

†Eogale
†Gracilocyon^{(paraphyletic genus)}
†Oodectes^{(paraphyletic genus)}
†Paramiacis
†Paroodectes

Incertae sedis	†"Miacis" sp. (CM 67873 &CM 77299)

Clade "C"

†Dormaalocyon
†Vulpavus clade	†Palaearctonyx †Vassacyon †Vulpavus^{(paraphyletic genus)} (including subgenus †Phlaodectes)
Incertae sedis	†"Miacis" deutschi †"Miacis" exiguus

Clade "D"

Order:Carnivora
(i.e. Clade "E")

Suborder:Caniformia	(SeeCaniformia)
Suborder:Feliformia	(SeeFeliformia)

Incertae sedis

other
representatives

†Africtis
†Dawsonicyon
†Miacis

Incertae sedis	†"Miacis" boqinghensis †"Miacis" hookwayi †"Miacis" latidens †"Miacis" petilus †Carnivoraformes undet. Genus A †Carnivoraformes undet. Genus B
ichnotaxa of Carnivoraformes	†Falcatipes

Superfamily:
†Viverravoidea

Family:
†Viverravidae

Subfamily:
†Didymictinae

†Bryanictis
†Didymictis
†Intyrictis
†Pristinictis
†Protictis^{(paraphyletic genus)} (including subgenus †Protictoides)
†Raphictis

Incertae sedis	†"Deltatherium" durini

Subfamily:
†Ictidopappinae

†Ictidopappus

Subfamily:
†Viverravinae

Incertae sedis

†Viverravidae sp. (CM 71188 &CM 71189)

Incertae sedis

†Hyaenodonta

Order:
†Hyaenodonta

Superfamily:
†Hyaenodontoidea

Family:
†Hyaenodontidae

†Cynohyaenodon/Quercytherium
clade

†Eurotherium clade

†Alienetherium
†Cartierodon
†Eurotherium^{(paraphyletic genus)}
†Paenoxyaenoides
†Prodissopsalis

†Leonhardtina clade

†Leonhardtina

†Matthodon clade

†Matthodon

†Oxyaenoides clade

†Oxyaenoides

Subfamily:
†Hyaenodontinae

†Consobrinus †Propterodon^{(paraphyletic genus)}
Tribe: †Hyaenodontini	†Hyaenodon (including subgenera †Neohyaenodon^{(paraphyletic subgenus)} and †Protohyaenodon^{(paraphyletic subgenus)})
Incertae sedis	†"Pterodon" hyaenoides

Clade:
†Proviverrinae

†Minimovellentodon †Morlodon †Parvagula
†Allopterodon/Proviverra clade	†Allopterodon †Lesmesodon †Proviverra

†Afro‑Arabian
clade

Superfamily:
†Hyainailouroidea
^{(polyphyletic superfamily)}

Family:
†Hyainailouridae
^{(paraphyletic family)}

Subfamily:
†Apterodontinae

†Apterodon
†Quasiapterodon

Subfamily:
†Hyainailourinae
^{(paraphyletic subfamily)}

†Akhnatenavus clade

†Maocyon/Orienspterodon
clade

†Maocyon
†Orienspterodon

Tribe:
†Hyainailourini
^{(polyphyletic tribe)}

†Bastetodon
†Exiguodon
†Falcatodon
†Hyainailouros^{(polyphyletic genus)}
†Parapterodon
†Sectisodon
†Sekhmetops
†Sivapterodon

Subtribe:
†Isohyaenodontina
^{(polyphyletic subtribe)}

†Isohyaenodon^{(polyphyletic genus)}

†Pterodon
clade

†Kerberos
Subtribe: †Pterodontina	†Pterodon

Incertae sedis

†"Pterodon" sp. (DPC 5036)

Tribe:
†Leakitheriini

†Leakitherium

Tribe:
†Metapterodontini

†Metapterodon

Tribe:
†Paroxyaenini

†Paroxyaena

Incertae sedis

Family:
†Prionogalidae

†Namasector
†Prionogale

Family:
†Teratodontidae

Subfamily:
†Teratodontinae

†Anasinopa †Brychotherium †Masrasector^{(paraphyletic genus)} †Metasinopa
Tribe: †Dissopsalini	†Buhakia †Dissopsalis
Tribe: †Teratodontini	†Teratodon
Incertae sedis	†Teratodontinae sp. (BC 2’08) †Teratodontinae sp. (CBI-1-614) †Teratodontinae sp. (Locality BQ-2, Fayum, Egypt)

other
representatives

†Furodon
†Glibzegdouia

†Kyawdawia clade

†Kyawdawia
†Paratritemnodon

†Lahimia clade

†Parvavorodon

Clade:
†Boualitominae
^{(paraphyletic clade)}

†Boualitomus †Lahimia
Incertae sedis	†Boualitominae sp. (Locality BQ-2, Fayum, Egypt) †Boualitominae sp. (Quarry L-41, Fayum, Egypt)

†Tritemnodon clade

†Tritemnodon

Clade:
†Koholiinae

†Koholia

Family:
†Indohyaenodontidae

†Indohyaenodon
†Yarshea

other
representatives

†Eoproviverra/Tinerhodon
clade

†Eoproviverra
†Tinerhodon

†Galecyon clade

Clade:
†Arfiinae

†Arfia

Clade:
†Limnocyoninae

†Iridodon †Limnocyon^{(paraphyletic genus)} †Oxyaenodon †Prolaena †Prolimnocyon^{(paraphyletic genus)} †Thinocyon
Incertae sedis	†"Thinocyon" sichowensis

Clade:
†Sinopidae

†Acarictis

†Sinopa clade

†Prototomus^{(paraphyletic genus)} †Sinopa
Incertae sedis	†Sinopidae sp. (FMNH PM 59529)

ichnotaxa
of Hyaenodonta

†Creodontipus †Dischidodacylus †Sarcotherichnus
Ichnofamily: †Sarjeantipodidae	†Hyaenodontipus †Quiritipes †Sarjeantipes

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