Typicallynocturnal, the caracal is highly secretive and difficult to observe. It isterritorial, and lives mainly alone or in pairs. The caracal is acarnivore that typically preys upon birds,rodents, and other smallmammals. It can leap higher than 3.0 m (9.8 ft) and catch birds in midair. It stalks its prey until it is within 5 m (16 ft) of it, after which it runs it down and kills it with a bite to the throat or to the back of the neck. Both sexes becomesexually mature by the time they are one year old and breed throughout the year. Gestation lasts between two and three months, resulting in a litter of one to six kittens. Juveniles leave their mothers at the age of nine to ten months, though a few females stay back with their mothers. The average lifespan of captive caracals is nearly 16 years.
The name 'caracal' was proposed byGeorges Buffon in 1761 who referred to itsTurkish name 'Karrah-kulak' or 'Kara-coulac', meaning 'black ear'.[3] The caracal is also known asdesert lynx andPersian lynx.[4] The 'lynx' of the Greeks and Romans was most probably the caracal, and the name 'lynx' is sometimes still applied to it, but the present-daylynx proper is a separate genus.[5]
In the 19th and 20th centuries, several caracalspecimens were described and proposed assubspecies. Since 2017, three subspecies have been recognised asvalid:[8]
Southern caracal (C. c. caracal)(Schreber, 1776) – occurs inSouthern andEast Africa
Results of aphylogenetic study indicates that the caracal and theAfrican golden cat (Caracal aurata)diverged between 2.93 and 1.19 million years ago. These two species together with theserval (Leptailurus serval) form theCaracallineage, which diverged between 11.56 and 6.66 million years ago.[11][12] The ancestor of this lineage arrived in Africa between 8.5 and 5.6 million years ago.[13]
The relationship of the caracal is considered as follows:[11][12]
A close facial view of a caracal, with the typical tufted ears and the black facial markings
The caracal is a slender, moderately sized cat characterised by a robust build, a short face, longcanine teeth, tufted ears, and long legs. It reaches nearly 40–50 cm (16–20 in) at the shoulder. The tan, bushy tail extends to thehocks.[14][15] The caracal issexually dimorphic; the females are smaller than the males in most bodily parameters.[16]
The prominent facial features include the 4.5 cm (1.8 in) long black tufts on the ears, two black stripes from the forehead to the nose, the black outline of the mouth, the distinctive black facial markings, and the white patches surrounding the eyes and the mouth. The eyes appear to be narrowly open due to the lowered upper eyelid, probably an adaptation to shield the eyes from the sun's glare. The ear tufts may start drooping as the animal ages. The coat is uniformly reddish tan or sandy, though black caracals are also known. The underbelly and the insides of the legs are lighter, often with small reddish markings.[16] The fur, soft, short, and dense, grows coarser in the summer. Theground hairs are denser in winter than in summer. The length of theguard hairs can be up to 3 cm (1.2 in) long in winter, but shorten to 2 cm (0.79 in) in summer.[17] These features indicate the onset ofmoulting in the hot season, typically in October and November.[18] The hind legs are longer than the forelegs, so the body appears to be sloping downward from therump.[15][16]
Male caracals measure in head-to-body length 78–108 cm (31–43 in) and have 21–34 cm (8.3–13.4 in) long tails; 77 male caracals ranged in weight between 7.2 and 19 kg (16 and 42 lb). The head-to-body length of females is 71–103 cm (28–41 in) with a tail of 18–31.5 cm (7.1–12.4 in); 63 females ranged in weight between 7 and 15.9 kg (15 and 35 lb).[19]
The caracal is often confused with a lynx, as both cats have tufted ears. However, a notable point of difference between the two is thatLynx species are spotted and blotched, while the caracal shows no such markings on the coat.[16] TheAfrican golden cat has a similar build as the caracal's, but is darker and lacks the ear tufts. Thesympatric serval can be distinguished from the caracal by the former's lack of ear tufts, white spots behind the ears, spotted coat, longer legs, longer tail, and smaller footprints.[17][20]
The skull of the caracal is high and rounded, featuring largeauditory bullae, a well-developedsupraoccipital crest normal to thesagittal crest, and a strong lower jaw. The caracal has a total of 30 teeth; thedental formula is3.1.3.13.1.2.1. Thedeciduous dentition is3.1.23.1.2. The canines are up to 2 cm (0.79 in) long and sharp. The caracal lacks the second upperpremolars, and the uppermolars are diminutive.[18] The large paws have fourdigits in the hind legs and five in the fore legs.[21][17] The first digit of the fore leg remains above the ground and features thedewclaw. The sharp and retractile claws are larger but less curved in the hind legs.[17]
In Africa, the caracal is widely distributed south of theSahara, but considered rare in North Africa. In Asia, it occurs from theArabian Peninsula, Middle East,Turkmenistan,Uzbekistan to western India.[1] It inhabits forests,savannas, marshy lowlands,semideserts, andscrub forests, but prefers dry areas with low rainfall and availability of cover. In montane habitats such as in theEthiopian Highlands, it occurs up to an elevation of 3,000 m (9,800 ft).[17]
In south-westernTurkey, caracals are present in a wildlife reserve in the provinces ofAntalya andMuğla that is regularly used for recreational and agricultural activities.[25] However, they avoid humans and are active mostly at night.[26] In Uzbekistan, caracals were recorded only in the desert regions of theUstyurt Plateau andKyzylkum Desert. Between 2000 and 2017, 15 individuals were sighted alive, and at least 11 were killed by herders.[27]
The caracal is typicallynocturnal, though some activity may be observed during the day as well. However, the cat is so secretive and difficult to observe that its activity at daytime might easily go unnoticed.[18] A study in South Africa showed that caracals are most active when the air temperature drops below 20 °C (68 °F); activity typically ceases at higher temperatures.[37] A solitary cat, the caracal mainly occurs alone or in pairs; the only groups seen are of mothers with their offspring.[15] Females inoestrus temporarily pair with males. A territorial animal, the caracalmarks rocks and vegetation in its territory with urine and probably with dung, which is not covered with soil. Claw scratching is prominent, and dung middens are typically not formed.[17] InIsrael, males are found to have territories averaging 220 km2 (85 sq mi), while that of females averaged 57 km2 (22 sq mi). The male territories vary from 270–1,116 km2 (104–431 sq mi) in Saudi Arabia. InMountain Zebra National Park, thehome ranges of females vary between 4.0 and 6.5 km2 (1.5 and 2.5 sq mi). These territories overlap extensively.[16] The conspicuous ear tufts and the facial markings often serve as a method of visual communication; caracals have been observed interacting with each other by moving the head from side to side so that the tufts flicker rapidly. Like other cats, the caracal meows, growls, hisses, spits, and purrs.[15]
In South Africa, caracals prey onCape grysbok,common duiker,bush vlei rats,rock hyrax andCape hare.[40][41][42] In areas where sheep and goat are farmed, caracals have also been documented preying on small livestock, but this is only a small and seasonal portion of their diet when wild prey is scarce.[40][19] Caracals are estimated to have caused about 11% ofAfrican penguin mortality inSimon's Town breeding colony between January 1999 and September 2021.[43]
The caracals's speed and agility make it an efficient hunter, able to take down prey two to three times its size.[1] The powerful hind legs allow it to leap more than 3 m (9.8 ft) in the air to catch birds on the wing.[16][44][45] It can even twist and change its direction mid-air.[16] It is an adroit climber.[16] It stalks its prey until it is within 5 m (16 ft), following which it can launch into a sprint. While large prey such as antelopes are suffocated by a throat bite, smaller prey are killed by a bite on the back of the neck.[16] Kills are consumed immediately, and less commonly dragged to cover. It returns to large kills if undisturbed.[17] It has been observed to begin feeding on antelope kills at the hind parts.[18] It mayscavenge at times, though this has not been frequently observed.[40]
Both sexes becomesexually mature by the time they are a year old; production ofgametes begins even earlier at seven to ten months. However, successful mating takes place only at 12 to 15 months. Breeding takes place throughout the year. Oestrus, one to three days long, recurs every two weeks unless the female is pregnant. Females in oestrus show a spike in urine-marking, and form temporary pairs with males. Mating has not been extensively studied; a limited number of observations suggest thatcopulation, lasting nearly four minutes on an average, begins with the male smelling the areas urine-marked by the female, which then rolls on the ground. He then approaches and mounts her. The pair separate after copulation.[16][17]
Gestation lasts about two to three months, following which a litter consisting of one to six kittens is born. Births generally peak from October to February. Births take place in dense vegetation or deserted burrows ofaardvarks andporcupines. Kittens are born with their eyes and ears shut and the claws not retractable (unable to be drawn inside); the coat resembles that of adults, but the abdomen is spotted. Eyes open by ten days, but it takes longer for the vision to become normal. The ears become erect and the claws become retractable by the third or the fourth week. Around the same time, the kittens start roaming their birthplace, and start playing among themselves by the fifth or the sixth week. They begin taking solid food around the same time; they have to wait for nearly three months before they make their first kill. As the kittens start moving about by themselves, the mother starts shifting them every day. All themilk teeth appear in 50 days, andpermanent dentition is completed in 10 months. Juveniles begin dispersing at nine to ten months, though a few females stay back with their mothers. The average lifespan of the caracal in captivity is nearly 16 years.[16][21][46]
In the 1990s, a captive caracal spontaneously mated with a domesticcat in theMoscow Zoo, resulting in afelid hybrid offspring.[47]
The caracal is listed asLeast Concern on theIUCN Red List since 2002, as it is widely distributed in over 50 range countries, where the threats to caracal populations vary in extent.Habitat loss due toagricultural expansion, the building of roads and settlements is a major threat in all range countries. It is thought to be close to extinction in North Africa, critically endangered in Pakistan, endangered in Jordan, but stable in central and Southern Africa. Local people kill caracal to protect livestock, or in retaliation for its preying on small livestock. Additionally, it is threatened by hunting for thepet trade on the Arabian Peninsula. In Turkey and Iran, caracals are frequently killed in road accidents.[1]In Uzbekistan, the major threat to caracal is killing by herders in retaliation for livestock losses. Guarding techniques and sheds are inadequate to protect small livestock like goats and sheep from being attacked by predators. Additionally, similarly to Ethiopia, heavy-traffic roads crossing caracal habitat pose a potential threat for the species.[27]
African caracal populations are listed underCITES Appendix II, while Asian populations come underCITES Appendix I. Hunting of caracal is prohibited in Afghanistan, Algeria, Egypt, India, Iran, Israel, Jordan, Kazakhstan, Lebanon, Morocco, Pakistan, Syria, Tajikistan, Tunisia, Turkey, Turkmenistan, and Uzbekistan. Caracals occur in a number of protected areas across their range.[1]
In South Africa, the caracal is considered Least Concern, as it is widespread and adaptable to a variety of habitats. It is tolerant to human-dominated areas, although it has been persecuted for many decades. Farmers are encouraged to report sightings of caracals, both dead and alive, and livestock killed by caracals to the national Predation Management Information Centre.[48]
The Central Asian caracal population is listed asCritically Endangered in Uzbekistan since 2009, and in Kazakhstan since 2010.[27][49][50]
A cheetah and a caracal kept for coursing by Rajasthani royalty, c. 1890[51]
The caracal appears to have been religiously significant in theancient Egyptian culture, as it occurs in paintings and as bronze figurines; sculptures are thought to have guarded the tombs ofpharaohs.[52] Embalmed caracals have also been discovered.[53]
The caracal was esteemed for its ability to catch birds in flight and was used forcoursing byMughal emperors in India at least since theDelhi Sultanate.[35] Chinese emperors used caracals as gifts. In the 13th and the 14th centuries,Yuan dynasty rulers bought numerous caracals,cheetahs (Acinonyx jubatus) andtigers (Panthera tigris) from Muslim merchants in the western parts of the empire in return for gold, silver,copper cash and silk. According to theMing Shilu, the subsequentMing dynasty continued this practice. Until the 20th century, the caracal was used in hunts of Indian rulers to hunt small game, while the cheetah was used for larger game.[54] In those times, caracals were used to huntbustards,francolins, and othergame birds.[55] They were also placed in arenas with flocks of pigeons and people would bet on which caracal would kill the largest number of pigeons. This probably gave rise to the expression "to put the cat among the pigeons".[45] Itspelt was used for making fur coats.[21]
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