| Broiliellus Temporal range:Early Permian | |
|---|---|
 | |
| Restoration ofBroiliellus olsoni | |
Scientific classification | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Order: | †Temnospondyli |
| Family: | †Dissorophidae |
| Subfamily: | †Dissorophinae |
| Genus: | †Broiliellus Williston,1914 |
| Type species | |
| †Broiliellus texensis Williston, 1914 | |
| Species | |
Broiliellus is an extinctgenus ofdissorophoidtemnospondyl within thefamilyDissorophidae.Broiliellus is most closely related to the genusDissorophus, and both have been placed in the subfamilyDissorophinae.Broiliellus is known from five species from theEarly Permian: thetype species isBroiliellus texensis, and the other species areBroiliellus brevis,Broiliellus olsoni, Broiliellus arroyoensis, andBroiliellus reiszi. An additional species,Broiliellus novomexicanus, which was originally namedAspidosaurus novomexicanus, is now thought to fall outside the genus as a member of the subfamilyEucacopinae.[1]
Broiliellus was first named by American paleontologistS.W. Williston in 1914 based on two nearly complete skulls in articulation with postcranial material from the early Permian of Texas; this species was given the nameBroiliellus texensis, the genus name being for the German paleontologistFerdinand Broili, and the species name being for the geographic provenance.[2] The holotype is currently reposited in theField Museum. A second species,Broiliellus brevis, from the Archer City bonebed in Texas was named in 1964 by Canadian paleontologistRobert Carroll; the holotype of this species is currently reposited in theMuseum of Comparative Zoology.[3] Two more species were named in 1967 by American paleontologist Robert DeMar,Broiliellus olsoni andBroiliellus arroyoensis, both from Texas.[4] The most recent species to be described wasBroiliellus reiszi in 2013 by Canadian paleontologist Robert Holmes and colleagues.[5]
Thephylogenetic analysis of Schoch (2012) found thatBroiliellus was most closely related toDissorophus. However, the three species ofBroiliellus did not form their ownclade, but rather apolytomy (unresolved evolutionary relationship) withDissorophus.[1]
Holmes et al. (2013) named a new species ofBroiliellus,Broiliellus reiszi.[5] They performed two phylogenetic analyses using the data matrix of Schoch (2012). This analysis yielded more resolved topology and amonophyleticBroiliellus. Below is thecladogram from that analysis:
| Dissorophoidea |
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