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Bract

From Wikipedia, the free encyclopedia
(Redirected fromBracteole)
Modified or specialized leaf
"Involucre" redirects here. For the protective layer that encloses spores in fungi, seePeridium. For the indusium in ferns or fungi, seeSorus.
Papery (upper) and leafy bracts on hay rattle (Rhinanthus minor). All the "leaves" in this image are bracts.

Inbotany, abract is a modified or specializedleaf, especially one associated with a reproductive structure such as aflower,inflorescence axis, orcone scale.

Bracts are usually different from foliage leaves; they may be of a different size, color, shape, or texture. Typically, they also look different from the parts of the flower, such as the petals orsepals.

A plant having bracts is referred to asbracteate[1] orbracteolate, while one that lacks them is referred to asebracteate[2] orebracteolate.

Variants

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Some bracts are brightly coloured and serve the function of attracting pollinators, either together with theperianth or instead of it. Examples of this type of bract include those ofEuphorbia pulcherrima (poinsettia) andBougainvillea: both of these have large colourful bracts surrounding much smaller, less colourful flowers.[3]

Ingrasses, each floret (flower) is enclosed in a pair of papery bracts, called thelemma (lower bract) andpalea (upper bract), while each spikelet (group of florets) has a further pair of bracts at its base calledglumes. These bracts form thechaff removed fromcereal grain duringthreshing andwinnowing.[4]

Bats may detect acoustic signals from dish-shaped bracts such as those ofMarcgravia evenia.[5]

Aprophyll is a leaf-like structure, such as a bracteole, subtending (extending under) a single flower orpedicel. The term can also mean the lower bract on apeduncle.

The frequently showy pair of bracts ofEuphorbia species in subgenusLacanthis are thecyathophylls.

Bracts subtend the cone scales in the seed cones of manyconifers, and in some cases, such asPseudotsuga, they extend beyond the cone scales.

Bracteole

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A small bract is called abracteole orbractlet. Technically this is any bract that ariseson a pedicel instead of subtending it.

Involucral bracts

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Beggar-tick (Bidens comosa)

Bracts that appear in awhorl subtending aninflorescence are collectively called aninvolucre. An involucre is a common feature beneath the inflorescences of manyApiaceae,Asteraceae,Dipsacaceae andPolygonaceae. Each flower in an inflorescence may have its own whorl of bracts, in this case called aninvolucel. In this case they may be calledchaff,paleas, orreceptacular bracts and are usually minute scales or bristles. Manyasteraceous plants have bracts at the base of each inflorescence.[6]

The term involucre is also used for a highly conspicuous bract or bract pair at the base of an inflorescence. In the familyBetulaceae, notably in the generaCarpinus andCorylus, theinvolucre is a leafy structure that protects the developing nuts. Beggar-tick (Bidens comosa) has narrow involucral bracts surrounding each inflorescence, each of which also has a single bract below it. There is then a pair of leafy bracts on the main stem and below those a pair of leaves.[6]

Epicalyx

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Epicalyx ofHibiscus rosa-sinensis

An epicalyx, which forms an additional whorl around thecalyx of a single flower, is a modification of bracteoles[7] In other words, the epicalyx is agroup of bracts resembling a calyx orbracteoles forming a whorl outer to the calyx.[8] It is a calyx-like extra whorl of floral appendages. Each individual segment of the epicalyx is called anepisepal because they resemble the sepals in them.[9] They are present in the hibiscus family,Malvaceae.Fragaria (strawberries) may or may not have an epicalyx.

Spathe

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Anthurium scherzerianuminflorescence with spathe andspadix

Aspathe is a large bract or pair of bracts forming a sheath to enclose the flower cluster of such plants aspalms,arums,irises,[10]crocuses,[11] anddayflowers (Commelina).Zephyranthes tubispatha in theAmaryllidaceae derives itsspecific name from its tubular spathe. In many arums (familyAraceae), the spathe is petal-like, attracting pollinators to the flowers arranged on a type of spike called aspadix.

See also

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References

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Wikimedia Commons has media related toBracts.
  1. ^"the definition of bracteate".Dictionary.com.Archived from the original on 2017-04-28. Retrieved2017-04-27.
  2. ^"Definition of Ebractate".www.greengonzo.com.Archived from the original on 2017-04-28. Retrieved2017-04-27.
  3. ^Environment Southwest. San Diego Society of Natural History. 1986.
  4. ^Jensen, William August; Salisbury, Frank B. (1972).Botany: an Ecological Approach. Wadsworth Publishing Company. p. 544.ISBN 978-0-534-00092-9.
  5. ^Ralph Simon; Marc W. Holderied; Corinna U. Koch; Otto von Helversen (2011). "Floral acoustics: Conspicuous echoes of a dish-shaped leaf attract bat pollinators".Science.333 (6042):631–633.Bibcode:2011Sci...333..631S.doi:10.1126/science.1204210.PMID 21798950.S2CID 5035286.
  6. ^abPopay, I (2022-01-07). Bidens frondosa (beggarticks) (Report).doi:10.1079/cabicompendium.108916.
  7. ^Darpan, Pratiyogita (June 2006).Competition Science Vision. Pratiyogita Darpan. p. 136.
  8. ^"Botany"(PDF). Archived fromthe original(PDF) on 2010-12-14. Retrieved2010-08-15.
  9. ^"epicalyx – Dictionary of botany". Botanydictionary.org.Archived from the original on 2012-05-01. Retrieved2012-04-29.
  10. ^Richard LynchThe Book of the Iris, p. 203, atGoogle Books
  11. ^Alex CashaFlora of the Maltese Islands, p. 287, atGoogle Books
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