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Borophagina

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Extinct subtribe of carnivores

Borophaginans Temporal range:Miocene–Pliocene PreꞒ Ꞓ O S D C P T J K Pg N
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Carnivora
Family:	Canidae
Subfamily:	†Borophaginae
Tribe:	†Borophagini
Subtribe:	†Borophagina X. Wang, 1999
Genera
Paratomarctus Carpocyon Protepicyon Epicyon Borophagus

Borophagina is asubtribe of theBorophaginae, a group of extinctcanids. They inhabited much ofNorth America from theEarly Miocene to theZanclean stage of thePliocene, 20.6—3.6Mya, and existed for approximately17 million years.^[1]

Like some other borophagines, they were short-faced, heavy-jawed canids^[2]^[3] although the group included both omnivorous and hypercarnivorous species.^[4]

References

[edit]

^PaleoBiology Database: Borophagina: Age range and collections
^Nowak, R. M., Walker's Mammals of the World. Maryland, Johns Hopkins University Press (edited volume) II, 1991
^Phylogenetic systematics of the Borophaginae (Carnivora, Canidae). Bulletin of the AMNH; no. 243, Wang, Tedford, Taylor.
^Wang, Xiaoming; Tedford, Richard H. (2008).Dogs, Their Fossil Relatives and Evolutionary History. Columbia. p. 35.ISBN 978-0-231-13528-3.

ExtinctCanidae

KingdomAnimalia
PhylumChordata
ClassMammalia
Order:Carnivora
Suborder:Caniformia

Canidae

†Prohesperocyon

†Hesperocyoninae

Hesperocyon Mesocyon Cynodesmus Sunkahetanka Philotrox Enhydrocyon Paraenhydrocyon Caedocyon Ectopocynus
Osbornodon	O. brachypus O. fricki O. iamonensis O. renjiei O. sesnoni O. scitulus O. wangi

†Borophaginae

Phlaocyonini

Cynarctoides
Phlaocyon	P. achoros P. annectens P. latidens P. leucosteus P. mariae P. marslandensis P. minor P. multicuspus P. taylori P. yatkolai

Borophagini

Cynarctina

Paracynarctus
Cynarctus

Aelurodontina

Tomarctus
Aelurodon

Borophagina

Paratomarctus Carpocyon Protepicyon Epicyon
Borophagus	B. diversidens B. dudleyi B. hilli B. littoralis B. orc B. parvus B. pugnator B. secundus

Caninae	see below↓

Caninae

†Leptocyon
†Metalopex

Urocyon	†U. citrinus †U. galushai †U. minicephalus †U. progressus †U. webbi

Vulpini

†Ferrucyon †Prototocyon
Nyctereutes (raccoon dogs)	†N. abdeslami †N. donnezani †N. lockwoodi †N. megamastoides †N. sinensis †N. tingi †N. vinetorum
Vulpes (true foxes)	†V. hassani †V. odessana †V. praeglacialis †V. qiuzhudingi †V. riffautae †V. rooki †V. skinneri †V. stenognathus

Canini(true dogs)

†Canis gezi
†Nurocyon

Cerdocyonina
(zorro)

†Protocyon †Theriodictis
Speothos	†Pleistocene bush dog (S. pacivorus)
†Dusicyon	Falkland Islands wolf (D. australis) D. avus D. cultridens
Lycalopex (South American foxes)	†L. cultridens (?) †L. ensenadensis

Canina(wolf-like canids)	see below↓

Falkland Islands wolf (Dusicyon australis)

Canina(wolf-like canids)

Canina

†Eucyon	E. davisi E. debonisi E. ferox E. intrepidus E. khoikhoi E. kuta E. marinae E. minor E. monticinensis E. skinneri E. wokari E. zhoui
Lycaon	†L. magnus †L. sekowei
Dhole (Cuon alpinus)	†C. a. antiquus †C. a. caucasicus †European dhole (C. a. europaeus) †Tien Shan dhole (C. a. hesperius)

Canis

Coyote (C. latrans)	†Pleistocene coyote (C. l. orcutti)
Red wolf (C. rufus)	†Florida black wolf (C. r. floridanus) †Texas red wolf (C. r. rufus)

Wolf (C. lupus)

†Prehistoric

†Recently
extinct

Sicilian wolf (C. l. cristaldii) Hokkaido wolf (C. l. hattai) Japanese wolf (C. l. hodophilax) Austro-Hungarian wolf
Synonymized underC. l. arctos:Bernard's wolf (C. l. bernardi)
Synonymized underC. l. occidentalis:Kenai Peninsula wolf (C. l. alces) Manitoba wolf (C. l. griseoalbus)
Synonymized underC. l. nubilus:Newfoundland wolf (C. l. beothucus) Cascade mountain wolf (C. l. fuscus) Mogollon mountain wolf (C. l. mogollonensis) Texas wolf (C. l. monstrabilis) Southern Rocky Mountain wolf (C. l. youngi)

See also:Evolution of the wolf
Category:Extinct canids

Taxon identifiers
Borophagina	Wikidata:Q4946287 EoL:61119819 Paleobiology Database:83324 Taxonomicon:6442565

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