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Black grouse

From Wikipedia, the free encyclopedia
Species of bird

Black grouse
Male
Bird recorded inFife, Scotland
Scientific classificationEdit this classification
Kingdom:Animalia
Phylum:Chordata
Class:Aves
Order:Galliformes
Family:Phasianidae
Genus:Lyrurus
Species:
L. tetrix
Binomial name
Lyrurus tetrix
(Linnaeus, 1758)
Distribution of black grouse globally
Black grouse range[2]
Close up of distribution of black grouse in Europe
Distribution inEurope.[2]
Synonyms[3]
  • Tetrao tetrix

Theblack grouse (Lyrurus tetrix), also known asnorthern black grouse,Eurasian black grouse,blackgame orblackcock,[4] is a largebird in thegrouse family. It is asedentaryspecies, spanning across thePalearctic inmoorland andsteppe habitat when breeding, often near wooded areas. They will spend the winter perched in dense forests, feeding almost exclusively on the needles of conifers. The black grouse is one of two species of grouse in the genusLyrurus, the other being the lesser-knownCaucasian grouse.

The female is greyish-brown and has a cackling or warbling call. She takes all responsibility for nesting and caring for the chicks, as typical with mostgalliforms.

The black grouse's genome was sequenced in 2014.[5]

Taxonomy and naming

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The black grouse wasformally described by the Swedish naturalistCarl Linnaeus in 1758 in thetenth edition of hisSystema Naturae under thebinomial nameTetrao tetrix.[6] BothTetrao andtetrix come fromAncient Greek words referring to some form of game bird.[7] The black grouse is now placed in the genusLyrurus that was introduced in 1832 by the English naturalistWilliam Swainson.[8][9]

The male and female are sometimes referred to by theirfolk names,blackcock andgreyhen, respectively. These names first occur in the literature withJohn Ray in 1674.[10]Heathcock andHeathhen are also common names.[11]

Subspecies

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The black grouse has six recognized subspecies.[12] Black grouse populations differ slightly in size and coloration, with birds increasing in size further east of their range:[13]

Description and appearance

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The black grouse is a large bird with males measuring roughly around 60 centimetres (24 inches) in length and weighing 1,100–1,250 g (2 lb 7 oz – 2 lb 12 oz), sometimes up to 2,100 g (4 lb 10 oz), with females approximately 45 cm (18 in) and weighing 750–1,100 g (1 lb 10 oz – 2 lb 7 oz).[13] The cock's fancy plumage is predominantly black with deep-blue hues on his neck and back, which contrasts the white wingline and undertail coverts, as well as red bare skin above each eye. On the other hand, the hen is much drabber and cryptically colored to blend in easily with the dense undergrowth, especially when nesting. The black grouse, along with the Caucasian grouse, has long outerrectrices (tail feathers) that curl outward and arranged in a way it resembles the frame of a Greeklyre, hence the genus name,Lyrurus.

Black grouse cock displaying

Distribution and habitat

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Black grouse can be found on open habitats acrossEurope (Swiss-Italian-French Alps especially) fromGreat Britain throughScandinavia,Estonia and acrossRussia. Although believed to once to live in Ireland, it now no longer resides there. In Eastern Europe they can be found inHungary,Latvia,Lithuania,Poland,Belarus,Romania andUkraine. There is a population in theAlps, and isolated remnants inGermany,France,Belgium and theNetherlands.[14] It formerly occurred inDenmark, but theDanish Ornithological Society (DOF) has considered it extinct since 2001. The species disappeared from Bulgaria in the 19th century.[15] In Asia, a huge portion of their population can be found in Russia (particularly southernSiberia), though they also inhabit parts ofKazakhstan,Mongolia,China, and possiblyKorea.[16]

Black grouse are adapted to an extensive array of habitats across Eurasia, though most frequently utilize the transitionary zones between forests and open clearings, especiallysteppe,heathland,grassland andpasture when near agricultural fields. Depending on the season, they will overwinter in large flocks in dense forests, and feed primarily on the leaves and buds of coniferous and broadleaf trees, such asScots pine,Siberian larch,silver birch, andEurasian aspen. Throughout the spring and summer, they tend to favor open spaces to seek potential mates and raise broods, switching their diet to berries, shoots and stems ofcranberries,bog bilberries,myrtleberries, and otherVaccinium shrubs.[13] They avoid the most extreme of desert and polar regions.

Conservation status

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Although this species has declined throughout most of its range in westernEurope, it is not considered to be vulnerable globally due to the large population (global estimate is 15–40 million individuals) and slow rate of decline.[1] Its decline is due toloss of habitat, disturbance, predation byfoxes,crows, etc., and small populations gradually dying out.[citation needed]

The IUCN implemented a Black Grouse Action Plan 2007–2010. This has looked at local populations that are vulnerable to theextinction vortex. For example, in Styria, Austria.[citation needed]

In the United Kingdom black grouse are found in upland areas of north Wales, thePennines and most of Scotland. Best looked for on farmland and moorland with nearby forestry or scattered trees. They have traditionallek sites where the males display.[citation needed]

They have declined in some parts of theUK (especiallyEngland), having disappeared from many of their former haunts. They are nowextirpated inLancashire,Derbyshire,Exmoor,East Yorkshire,New Forest,Nottinghamshire,Worcestershire,Quantock Hills,Cornwall,Dartmoor,Kent,Wiltshire andSurrey.[citation needed]

A programme to re-introduce black grouse into the wild started in 2003 in theUpper Derwent Valley area of thePeak District inEngland. 30 grouse were released in October 2003, followed by 10 male grouse in December 2004 and a further 10 males and 10 females in April 2005. The programme is being run jointly by theNational Trust,Severn Trent Water andPeak District National Park.[citation needed]

Conservation groups helping to revive the black grouse include theRSPB and theGame & Wildlife Conservation Trust.[citation needed]

In France there has been much work regarding the birds and their risk of flying intoski-lifts.[17]

From 1950 to 2000, local black grouse populations have steadily diminished inManchuria and northeastern China by about 39%, with birds being most affected (and possibly extirpated) in theJilin Province. Though the exact causes for local declines remain largely unclear, habitat loss and excess hunting have played a part in the overall reduction. Shifts in their location, however can happen concurrently with the depletion over decades, so short-term research projects may not provide much proof of driving forces behind population alterations.[18]

Based on historical info gathered from various sources (including wildlife surveys and scientific papers), aspen, birch, and poplar make up the black grouse's staple diet and habitat. The climate and precipitation during the month of June is also linked to the breeding success of black grouse.[18]

Breeding and nesting

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Egg ofTetrao tetrix tetrix -MHNT
Black grouse - rolling and hissing sounds, recorded in 2022 in the northern Alps
Black grouse - recorded in 2022 in the northern Alps. Mostly rolling sounds audible.

Black grouse have a very distinctive and well-recorded courtship ritual. Every dawn in the spring, male grouse begin competition with other males in hopes of attracting a hen to mate with. They will display to signal their territory and vigor by fanning out their elaborate lyre-shaped tails and inflating their necks on designated open ground called alek. Their song consists of a long, dove-like bubbling coo or murmur. Black grouse hens visiting the lek decide the overall healthiest male, though not all females may arrive at every lek.

In western Europe, these leks seldom contain more than 40 birds; inRussia, 150 is not uncommon and 200 have been recorded.[14]

When mated successfully, she will fly away from the site to a suitable nesting site with an abundance of dense shrub or tall vegetation, often located at a tree base in between roots, under low branches, beside a boulder, or extremely rarely, a used raptor's or corvid's nest 7 metres (23 ft) off the ground. A dent (23–28 cm or 9–11 in wide by 10–11 cm or 4–4+12 in deep) is scraped out on the dirt floor and cushioned with grasses, sticks, leaves, and feathers. About 6–11 pale buff eggs speckled brown are then laid in the nest, incubated for approx. 23–28 days. The chicks consume invertebrates, transitioning to more plant matter as they mature. By around 10–14 days and so forth, they are capable of short flights.[13]

Where their range overlaps in similar biomes of other species, they are capable of hybridizing with theringneck pheasant,western capercaillie,black-billed capercaillie,Siberian grouse,hazel grouse, andwillow ptarmigan.[4]

Relationship to humans

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The black grouse in the coat of arms ofTuusniemi

The tails of black-cocks have, since late Victorian times, been popular adornments for hats worn withHighland Dress. Most commonly associated withGlengarry andBalmoral orTam o' Shanter caps, they still continue to be worn by pipers of civilian and militarypipe bands. Since 1904, all ranks of theRoyal Scots andKing's Own Scottish Borderers have worn them in their full-dress headgear and that tradition is carried on in the dress glengarries of the current Scottish super regiment, theRoyal Regiment of Scotland.[citation needed]

Gallery

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  • Male displaying
    Male displaying
  • Painting of a male and female
    Painting of a male and female
  • Female
    Female
  • On snow
    On snow
  • Taxidermies with different color anomalies
    Taxidermies with different color anomalies

References

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  1. ^abBirdLife International (2016)."Lyrurus tetrix".IUCN Red List of Threatened Species.2016 e.T22679480A85944601.doi:10.2305/IUCN.UK.2016-3.RLTS.T22679480A85944601.en. Retrieved12 November 2021.
  2. ^abBirdLife International and NatureServe (2014) Bird Species Distribution Maps of the World. 2012. Lyrurus tetrix. In: IUCN 2014. The IUCN Red List of Threatened Species. Version 2014.3."The IUCN Red List of Threatened Species". Archived fromthe original on 27 June 2014. Retrieved27 June 2014.. Downloaded on 2 June 2015.
  3. ^"Lyrurus tetrix".Avibase.
  4. ^ab"Lyrurus tetrix (Black Grouse) - Avibase".avibase.bsc-eoc.org. Retrieved1 October 2020.
  5. ^Wang, B.; Ekblom, R.; Bunikis, I.; Siitari, H.; Höglund, J. (6 March 2014)."Whole genome sequencing of the black grouse (Tetrao tetrix): Reference guided assembly suggests faster-Z and MHC evolution".BMC Genomics.15 (1): e10000180.doi:10.1186/1471-2164-15-180.PMC 4022176.PMID 24602261.
  6. ^Linnaeus, Carl (1758).Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis (in Latin). Vol. 1 (10th ed.). Holmiae (Stockholm): Laurentii Salvii. p. 159.
  7. ^Jobling, James A (2010).The Helm Dictionary of Scientific Bird Names. London: Christopher Helm. p. 383.ISBN 978-1-4081-2501-4.
  8. ^Swainson, William;Richardson, J. (1831).Fauna Boreali-Americana, or, The Zoology of the Northern Parts of British America. Vol. Part 2. The Birds. London: J. Murray. p. 497. The title page bears the year 1831 but the volume was no published until 1832.
  9. ^Gill, Frank; Donsker, David;Rasmussen, Pamela, eds. (July 2021)."Pheasants, partridges, francolins".IOC World Bird List Version 11.2. International Ornithologists' Union. Retrieved23 August 2021.
  10. ^Lockwood, W.B. (1993).The Oxford Dictionary of British Bird Names.OUP.ISBN 978-0-19-866196-2.
  11. ^Rines, George Edwin, ed. (1920)."Blackcock" .Encyclopedia Americana.
  12. ^"IOC World Bird List 13.1".IOC World Bird List.doi:10.14344/ioc.ml.13.1.
  13. ^abcdde Juana, Eduardo; Kirwan, Guy M.; Boesman, Peter F. D. (4 March 2020)."Black Grouse (Tetrao tetrix)".Birds of the World.doi:10.2173/bow.blagro1.01.1.S2CID 243567408.
  14. ^abThe Birds of the Western Palearctic (Abridged ed.).OUP. 1997.ISBN 0-19-854099-X.
  15. ^Boev, Z. (1997). "The Black Grouse, Tetrao tetrix (L., 1758) (Tetraonidae, Aves), a disappeared species in Bulgaria (Paleolithic and Neolithic records)".Anthropozoologica.25–26:643–646.
  16. ^"Black Grouse - eBird".ebird.org. Retrieved25 September 2020.
  17. ^"Observatoire des galliformes de montagne (Observatory of Mountain Galliformes)" (in French). Archived fromthe original on 3 December 2013. Retrieved25 November 2013.
  18. ^abZhang, Chao; Yang, Li; Wu, Shuhong; Xia, Wancai; Yang, Lei; Li, Miaomiao; Chen, Minhao; Luan, Xiaofeng (2020)."Use of historical data to improve conservation of the black grouse (Lyrurus tetrix) in Northeast China".Ecosphere.11 (3): e03090.Bibcode:2020Ecosp..11E3090Z.doi:10.1002/ecs2.3090.ISSN 2150-8925.

External links

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Game animals andshooting in the United Kingdom
Game birds
Quarry species
Deer
Other quarry
Opponents
Law
See also
1 Rarely shot because of declining numbers.
Lyrurus tetrix
Tetrao tetrix
International
National
Other
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