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| Bitternut hickory | |
|---|---|
| Bitternut hickory foliage | |
| Scientific classification | |
| Kingdom: | Plantae |
| Clade: | Tracheophytes |
| Clade: | Angiosperms |
| Clade: | Eudicots |
| Clade: | Rosids |
| Order: | Fagales |
| Family: | Juglandaceae |
| Genus: | Carya |
| Section: | Carya sect.Apocarya |
| Species: | C. cordiformis |
| Binomial name | |
| Carya cordiformis | |
| Natural range | |
Carya cordiformis, thebitternut hickory,[2] also called bitternut, yellowbud hickory, or swamp hickory, is a large hickory species native to the eastern United States and adjacent Canada. Notable for its unique sulphur-yellow buds, it is one of the most widespread hickories and is the northernmost species of pecan hickory (Carya sect.Apocarya). It is the shortest-lived of the hickories, living to about 200 years.[3]
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It is a largedeciduoustree, growing up to 35 m (115 ft) tall (exceptionally to 47 m or 154 ft), with a trunk up to 1 m (3 ft 3 in) diameter. Theleaves are 15–30 cm (6–12 in) long, pinnate, with 7–11 leaflets, each leaflet lanceolate, 7–13 cm (2+3⁄4–5 in) long, with the apical leaflets the largest but only slightly so. Theflowers are small wind-pollinatedcatkins, produced in spring. Thefruit is a very bitternut,2–3 cm (3⁄4–1+1⁄4 in) long with a green four-valved cover which splits off at maturity in the fall, and a hard, bony shell. Another identifying characteristic is its bright sulfur-yellow winter bud.
It is closely related to thepecan, sharing similar leaf shape and being classified in the same section of the genusCarya sect.Apocarya, but unlike the pecan, it does not have edible nuts. It is most readily distinguished from the pecan by the smaller number of leaflets, with many leaves having only 7 leaflets (rarely fewer than 9, and often 11–13, in the pecan).Hybrids with the pecan are known, and namedCarya ×brownii. A hybrid between the shagbark hickory (C. ovata) is also recognized, and is known as Laney's hickory (Carya ×laneyi).[4]
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Bitternut hickory grows in moist mountain valleys along streambanks and in swamps. Although it is usually found on wet bottom lands, it grows on dry sites and also grows well on poor soils low in nutrients. The species is not included as a titled species in the Society of American Foresters forest cover types because it does not grow in sufficient numbers.
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Bitternut hickory grows throughout the eastern United States from southwestern New Hampshire, Vermont, Maine, and southern Quebec; west to southern Ontario, central Michigan, and northern Minnesota; south to eastern Texas; and east to northwestern Florida and Georgia. It is most common, however, from southern New England west to Iowa and from southern Michigan south to Kentucky. It is probably the most abundant and most uniformly distributed of all the hickories. It is most commonly found in the Southern Appalachian Mountains, in high elevations.[5]
The taxon was first described in 1787 asJuglans cordiformis byFriedrich Adam Julius von Wangenheim,[6][7] and was transferred to the genus,Carya, in 1869 byKarl Koch.[6][8]
Bitternut is used for lumber and pulpwood. Commercial stands are located mostly north of the other pecan hickories. Bitternut hickory is cut and sold in mixture with the "true" hickories. Because bitternut hickory wood is hard and durable, it is used for furniture, paneling, dowels, tool handles and ladders. Like other hickories, thewood is used forsmokingmeat, and byNative Americans for makingbows. Bitternut hickory seeds are eaten by rabbits,[9] and both its seeds and bark are eaten by other wildlife.
Thetannins which give the nuts of bitternut hickory their bitter flavour are notfat soluble. As a result, it is possible to extract an edible oil from the nuts through pressing and separating the tannin containing pulp from the oil.[10] Reportedly, the oil content of bitternut hickories reaches as high as 80%.[11] The oil of bitternut hickory is reputed to have a mild and pleasant flavour, similar to that ofpecan oil.[10]
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Bitternut hickory is adiploid species with two sets of sixteen chromosomes that readily hybridizes with other diploid hickory species with a few namedhican varieties available. The pecan variety 'Major' has bitternut alleles at twosimple sequence repeat loci indicating a cryptic cross that may also have involvedC. ovata.