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Arandaspida

From Wikipedia, the free encyclopedia
Extinct subclass of jawless fishes

Arandaspida
Life restoration ofArandaspis
Scientific classificationEdit this classification
Domain:Eukaryota
Kingdom:Animalia
Phylum:Chordata
Infraphylum:Agnatha
Class:Pteraspidomorpha
Subclass:Arandaspida
Ritchie and Gilbert-Tomlinson, 1977
Families
Sacabambaspis, the best known arandaspid, from the Ordovician of Bolivia. shows the characteristic, frontally positioned eyes, like car head lamps[1]

Arandaspida is a taxon of very early, jawlessprehistoric fish which lived during theOrdovicianperiod. Arandaspids represent some of the oldest knownvertebrates. The group represents asubclass within the classPteraspidomorphi, and contains only one order, theArandaspidiformes. The oldest known genus of this group isSacabambaspis found inSouth America.

Characteristics

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The head armor of arandaspids is elongated, fusiform, with a rather flat dorsal shield, and a bulging ventral shield. In the anterior part of the dorsal shield are two closely set holes, which have been thought to be a paired pineal opening, but which are more likely the external openings of the endolymphatic ducts.

The eyes, surrounded by a sclerotic ring, are housed in a notch at the anterior end of the dorsal shield. The nostrils are not clearly located, but may have been situated between the eyes. Ventrally, the ventral lip of the mouth is armed with long series of small oral plates which recall those ofheterostracans.

The gill openings are probably numerous (more than 15) and minute. They opened between the diamond-shaped platelets which separate the dorsal from the ventral shield.

The body is covered with rod-shaped scales arranged in chevrons, and the tail is probably pad-shaped and diphycercal. The dermal bones of arandaspids consist of aspidine (acellular bone) and are ornamented with oakleaf-shaped tubercles which seem to contain no dentine. The sensory-lines were housed in narrow grooves between the tubercles.

Taxonomy

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Taxonomy based on the work of Mikko's Phylogeny Archive,[2] Nelson, Grande and Wilson 2016[3] and van der Laan 2018.[4]

  • Order †ArandaspidiformesRitchie & Gilbert-Tomlinson, 1977
    • FamilyAstraspididaeHalstead 1993
    • FamilyArandaspididaeRitchie & Gilbert-Tomlinson, 1977 [PorophoraspididaeHalstead 1993; Sacabambaspidae]

See also

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References

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  1. ^Janvier, Philippe (1997)ArandaspidaThe Tree of Life Web Project.
  2. ^Haaramo, Mikko (2003)."Pteraspidomorphi".in Mikko's Phylogeny Archive. After Carroll, 1988, and Janvier, 1997. Retrieved26 October 2013.
  3. ^Nelson, Joseph S.; Grande, Terry C.; Wilson, Mark V. H. (2016).Fishes of the World (5th ed.).John Wiley & Sons.ISBN 9781118342336.
  4. ^van der Laan, Richard (2017)."Family-group names of fossil fishes".{{cite journal}}:Cite journal requires|journal= (help)
Arandaspida
Arandaspidiformes
Astraspididae
Arandaspididae
Heterostracomorphi
Astraspida
Astraspidiformes
Astraspididae
Tesakoviaspidida
Tesakoviaspididae
Eriptychiida
Eriptychiiformes
Eriptychiidae
Eriptychiidae
Oniscolepididae
Heterostraci
    • see below↓
ArandaspisAstraspis desiderata
Cyathaspidiformes
    • see below↓
Pteraspidiformes
    • see below↓
Cardipeltida
Lepidaspidida
Corvaspidida
Tolypelepidida
Tesseraspidiformes
Traquairaspidiformes
Phialaspididae
Traquairaspididae
Weigeltaspididae
Athenaegis chattertoniHeterostraci
Cyathaspidida
Cyathaspidae
Ariaspidae
Ctenaspidae
Amphiaspidida
incertae sedis
Amphiaspidoidei
Amphiaspididae
Edaphaspididae
Gabreyaspididae
Olbiaspididae
Hibernaspidoidei
Hibernaspididae
Eglonaspididae
Aphataspididae
Siberiaspidoidei
Siberiaspididae
Tuxeraspididae
PoraspisHibernaspis macrolepis
Anchipteraspididae
Podolaspididae
Protopteraspididae
Psammosteida
Drepanaspididae
Guerichosteidae
Pycnosteidae
Psammolepididae
Psammosteidae
Obrucheviidae
Pteraspidoidei
incertae sedis
Pteraspididae
Gigantaspididae
Protaspididae
Pteraspis rostrataPsammolepis alata
Arandaspida
Arandaspidiformes
Arandaspidida


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