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Arabian leopard

From Wikipedia, the free encyclopedia
Subspecies of mammals

Arabian leopard
Leopard atEin Gedi,Israel
Scientific classificationEdit this classification
Kingdom:Animalia
Phylum:Chordata
Class:Mammalia
Order:Carnivora
Family:Felidae
Genus:Panthera
Species:
Subspecies:
P. p. nimr
Trinomial name
Panthera pardus nimr
(Hemprich andEhrenberg), 1833
Distribution of the Arabian leopard
Synonyms

P. p. jarvisiPocock, 1932

TheArabian leopard (Panthera pardus nimr) is the smallestleopardsubspecies. It was described in 1830 and is native to theArabian Peninsula, where it was widely distributed in rugged hilly and montane terrain until the late 1970s. Today, the population is severely fragmented and thought to decline continuously. In 2008, an estimated 45–200 individuals in three isolated subpopulations were restricted to westernSaudi Arabia,Oman andYemen. However, as of 2023, it is estimated that 100–120 in total remain, with 70-84 mature individuals, in Oman and Yemen, and it is possibly extinct in Saudi Arabia. The current population trend is suspected to be decreasing.[1]

Taxonomic history

[edit]

Felis pardus nimr was thescientific name proposed byWilhelm Hemprich andChristian Gottfried Ehrenberg in 1830 for a leopard from Arabia.[2]Panthera pardus jarvisi, proposed byReginald Innes Pocock in 1932, was based on a leopard skin from theSinai Peninsula.[3]

In the early 1990s, aphylogeographic analysis was carried out based on tissue samples from Asian andAfrican leopards.P. p. jarvisi was provisionally grouped withPanthera pardus tulliana, as tissue samples were not available.[4]Genetic analysis of a single wild leopard fromSouth Arabia appeared most closely related to the African leopard, andMolecular biologists tentatively proposed in 2001 to group the Sinai leopard with the Arabian leopard, as again tissue samples were not available.[5]

Characteristics

[edit]

The Arabian leopard's fur varies from pale yellow to deep golden, tawny or grey and is patterned withrosettes.[6] Males have a total length of 182–203 cm (72–80 in) including 77–85 cm (30–33 in) long tails and weigh about 30 kg (66 lb); females are 160–192 cm (63–76 in) long including 67–79 cm (26–31 in) long tails and weigh around 20 kg (44 lb).[7] It is the smallest leopard subspecies.[8]It is however the largest cat in the Arabian Peninsula.[9][10]

Distribution and habitat

[edit]
Captive leopard in theBreeding Centre for Endangered Wildlife, Sharjah

The geographic range of the Arabian leopard is poorly understood but generally considered to be limited to the Arabian Peninsula, includingEgypt'sSinai Peninsula.[11] It lives in mountainous uplands and hilly steppes, but seldom moves to open plains, desert or coastal lowlands.[10] Since the late 1990s, leopards were not recorded in Egypt.[7] One individual was killed in theElba Protected Area in 2014.[12]

Until the late 1960s, the Arabian leopard was widely distributed in the mountains along both the coasts of theRed Sea andArabian Sea.[9] InSaudi Arabia, leopard habitat is estimated to have decreased by around 90% since the beginning of the 19th century. Of 19 reports obtained from informants between 1998 and 2003, only four are confirmed including sightings in one location in theHijaz Mountains and three locations in theAsir Mountains, with the most recent record in 2002 south ofBiljurashi. No leopard was recorded during acamera trapping survey conducted from 2002 to 2003. Although the leopard is officially protected in the country, its remaining range is not encompassed by protected areas.[13]

In theUnited Arab Emirates, the Arabian leopard was first sighted in 1949 byWilfred Thesiger inJebel Hafeet.[14] The exact status of the leopard in the country is unclear. It is eitherextinct or very rare in the eastern region, with occasional sightings being reported in places likeWadi Wurayah.[15] Before the end of the 20th century, sightings were reported in the areas of Jebel Hafeet and Al-Hajar Mountains.[7][16]

InOman, leopards were reported to have occurred in the Hajar Mountains until the late 1970s.[7] The largest confirmed sub-population inhabits theDhofar Mountains in the country'ssoutheast. In theJabal Samhan Nature Reserve, 17 individual adult leopards were identified between 1997 and 2000 using camera traps.[17] Leopards were also sighted in theMusandam Peninsula,[7] particularlyRas Musandam.[9] The home range of Arabian leopards in this reserve is roughly estimated at 350 km2 (140 sq mi) for males and 250 km2 (97 sq mi) for females.[16] The Dhofar mountain range is considered the besthabitat for leopards in the country. This rugged terrain provides shelters, shade and trapped water, and harbors a wide variety of prey species, in particular in escarpments and narrowwadis.[18]

InYemen, leopards formerly ranged in all mountainous areas of the country, including the western and southern highlands eastwards to the border with Oman. Since the early 1990s, leopards are considered rare and close to extinction due to direct persecution by local people and depletion of wild prey.[19]

There was a small population inIsrael'sNegev desert, estimated at 20 individuals in the late 1970s.[20] Leopards were hunted until the early 1960s. By 2002, fewer than 11 isolated individuals were estimated to survive. Six males, three females and two unsexed individuals were identified in the country, based on genetic analysis of 268 scats collected. About five individuals were thought to survive in theJudaean Desert as of 2005.[21] The last wild leopard in the Negev desert was sighted nearSde Boker in 2007, which was in a poor and weak shape; and the last leopard in the northernArabah Valley was sighted in 2010–11.[22]

InJordan, the last confirmed sighting of a leopard dates to 1987.[23]

Ecology and behaviour

[edit]
A South Arabian relief from the 5th century BC, inWalters Art Museum

Arabian leopards are predominantlynocturnal, but are sometimes also seen in daylight.[10] They seem to concentrate on small to mediumprey species, and usually store carcasses of large prey in caves or lairs but not in trees.[24] Scat analyses revealed that the main prey species compriseArabian gazelle,Nubian ibex,Cape hare,rock hyrax,porcupine,Ethiopian hedgehog, smallrodents, birds, andinsects. Since local people reducedungulates to small populations, leopards are forced to alter their diet to smaller prey andlivestock such as goats, sheep, donkeys and young camels.[13]

Information about ecology and behaviour of Arabian leopards in the wild is very limited.[16] A leopard from the Judean desert is reported to have come into heat in March. After agestation period of 13 weeks, females give birth to two to four cubs in a cave amidst boulders or in a burrow.[10]

Leopard cubs are born with closed eyes that open four to nine days later.[25] Captive-born Arabian leopard cubs emerged from their den for the first time at the age of one month.[26] Cubs areweaned at the age of about three months, and remain with their mother for up to two years.[25]

Threats

[edit]
Stuffed leopard from theSinai Peninsula in theGiza Zoological Museum,Egypt
Taxidermied Arabian leopard at theOman Natural History Museum inMuscat

Three confirmed separate subpopulations remain on the Arabian Peninsula with fewer than an estimated 200 leopards.[27][28] The Arabian leopard is threatened by habitat loss,degradation and fragmentation; prey depletion caused by unregulated hunting; trapping for theillegal wildlife trade and retaliatory killing in defense of livestock.[7][13]

The leopard population has decreased drastically in Arabia as shepherds and villagers kill leopards in retaliation for attacks onlivestock. In addition, hunting of leopard prey species such ashyrax andibex by local people andhabitat fragmentation, especially in theSarawat Mountains, made the continued survival of the leopard population uncertain. Other reasons for killing leopards are for personal satisfaction and pride, traditionalmedicine and hides. Some leopards are killed accidentally when eating poisoned carcasses intended forArabian wolf andstriped hyena.[29]Among the products sold in the tent city ofMina, Saudi Arabia after theHajj of 2010, skins of Arabian leopards that were poached in Yemen were offered.[30]

The leopard population in Saudi Arabia is affected by the decrease of natural prey species so that leopards increasingly prey on livestock. Local people therefore consider leopards a threat and kill them either by using poison or snares. The leopard population is close to extinction in the country.[31]

TheIsraeli West Bank Barrier, built by the Israeli government in the mid-2000s, disrupts themigration of all terrestrial land animals between Israel and Palestine, therefore reducing animals' natural habitats and causing gene pools to be significantly shallower, which results ininbreeding andinfertility.[32]

In the 1950s, the Arabian leopard population was already decreasing drastically due to habitat degradation and fragmentation, and killing of leopards and prey species.[33]

Conservation

[edit]

The 4,500 km2 (1,700 sq mi) Jabal Samhan Nature Reserve was established in 1997 aftercamera trap records of leopards were obtained; in the following decade, 17 individual adult leopards and one cub were identified.[17] Leopards were also radio-collared and tracked in this Nature Reserve.[34]

At least ten wild leopards were live-captured in Yemen since the early 1990s and sold to zoos; some have been placed in conservation breeding centers in the UAE and Saudi Arabia.[19]

A detailed study of leopard distribution and habitat requirement is needed for the management of the species. The ecological information needed include data on feeding behavior, range use and reproduction. This information is of great importance to the survival of the species. There are many sites already surveyed and considered to be suitable for preservation for leopards in the plan adopted by the national commission for wildlife conservation and development. These areas include Jebel Fayfa, Jebel Al-Qahar, Jebel Shada, which has already been gazetted as a protected area, Jebel Nees, Jebel Wergan, Jebel Radwa and Harrat Uwayrid. The formal establishment of some of these areas is now urgent.[1]

A successfulconservation strategy must promote the awareness of the importance of leopard conservation, employing the media and perhaps other sources for basic education programs. The support and involvement of people living close to leopard habitats are vital in such efforts. This is true not only because they might affect the conservation of the leopard in one way or another, but also because they depend on their livestock which could be killed occasionally by leopards. Although it is not always practical, compensation for lost livestock from leopard predation should be considered.[35]

Revenue from sources such as hunting rights andecotourism, services such as roads and school employment in protected areas would encourage local residents to participate in leopard conservation. Furthermore, well-managed protected areas will ensure the continued survival of the species until other factors enhancing its survival become effective. Public awareness, fruitful consideration of the needs of local people and ecological studies may take years to be useful.[36]

In Yemen, efforts are underway to conserve leopards at two sites, including Hawf Protected Area.[37] In Saudi Arabia, authorities have undertaken efforts to create Sharaan Nature Reserve, a wildlife sanctuary for the leopard in the area ofAl-`Ula.[38][39]

In Israel, Arabian leopards are being bred inYotvata Hai-Bar Nature Reserve for future reintroduction.[40]

In captivity

[edit]
Leopard at the former Arabia's Wildlife Centre in Sharjah, the UAE

The first Arabian leopards were captured in southern Oman and registered in thestudbook in 1985.Captive breeding was initiated in 1995 at the Oman Mammal Breeding Centre and is operated at a regional level on the Arabian Peninsula. Since 1999, the regional studbook is coordinated and managed by personnel of theBreeding Centre for Endangered Arabian Wildlife, Sharjah.[41] As of 2010[update], nine institutions participated in the breeding programme and kept 42 males, 32 females, and three unsexed leopards, of which 19 were wild caught. This captive population comprised 14 founders that have an unequal number of descendants.[42] In 2016, the leopards and other fauna were transferred from the breeding centre in Sharjah toAl Hefaiyah Conservation Centre in the eastern area ofKalba.[43]

In Yemen, leopards were kept atTa'izz andSana'a Zoos.[44] Two cubs were born on 26 April 2019 at the Prince Saud Al-Faisal Wildlife Research Center inTa'if.[45]

Arabian Leopard Day

[edit]
Arabian Leopard Day Logo

In February 2022,Saudi Council of Ministers declaredFebruary 10 as the "Arabian Leopard Day" in an effort to protect the species and raise awareness of their conservation status.[46] In June 2023, TheUnited Nations voted to adopt a resolution to officially designate February 10 as an international day for Arabian Leopards.[47]

On the second Arabian Leopard Day, which took place in February 2023, TheRoyal Commission ofAl Ula created a $25 million fund to promote conservation efforts and signed a 10-year deal withPanthera worth $20 million.[48]

See also

[edit]
Leopard subspecies

References

[edit]
  1. ^abcAl Hikmani, H.; Spalton, A.; Zafar-ul Islam, M.; al-Johany, A.; Sulayem, M.; Al-Duais, M.; Almalki, A. (2024) [amended version of 2023 assessment]."Panthera pardus ssp.nimr".IUCN Red List of Threatened Species.2024 e.T15958A259040222.doi:10.2305/IUCN.UK.2024-1.RLTS.T15958A259040222.en.
  2. ^Hemprich, W.; Ehrenberg, C. G. (1830)."Felis, pardus?, nimr". In Dr. C. G. Ehrenberg (ed.).Symbolae Physicae, seu Icones et Descriptiones Mammalium quae ex Itinere per Africam Borealem et Asiam Occidentalem Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg. Decas Secunda. Zoologica I. Mammalia II. Berolini: Officina Academica. pp. Plate 17.
  3. ^Pocock, R. I. (1932). "The Leopards of Africa".Proceedings of the Zoological Society of London: 543−591.doi:10.1111/j.1096-3642.1932.tb01085.x.
  4. ^Miththapala, S.; Seidensticker, J.; O'Brien, S. J. (1996). "Phylogeographic Subspecies Recognition in Leopards (P. pardus): Molecular Genetic Variation".Conservation Biology.10 (4):1115–1132.doi:10.1046/j.1523-1739.1996.10041115.x.
  5. ^Uphyrkina, O.; Johnson, W. E.; Quigley, H.; Miquelle, D.; Marker, L.; Bush, M. & O'Brien, S. (2001)."Phylogenetics, genome diversity and origin of modern leopard,Panthera pardus"(PDF).Molecular Ecology.10 (11):2617–2633.Bibcode:2001MolEc..10.2617U.doi:10.1046/j.0962-1083.2001.01350.x.PMID 11883877.S2CID 304770. Archived fromthe original(PDF) on 2020-04-28. Retrieved2010-03-02.
  6. ^Seidensticker, J. & Lumpkin, S. (1991).Great Cats. London: Merehurst.
  7. ^abcdefSpalton, J. A.; Al Hikmani, H. M. (2006)."The Leopard in the Arabian Peninsula – Distribution and Subspecies Status"(PDF).Cat News (Special Issue 1):4–8. Archived from the original on May 23, 2011.
  8. ^Biquand, S. (1990). "Short review of the status of the Arabian leopard,Panthera pardus nimr, in the Arabian Peninsula". In Shoemaker, A. (ed.).International Leopard Studbook. Columbia: Riverbanks Zoological Park. pp. 8−10.
  9. ^abcNader, I. A. (1989)."Rare and endangered mammals of Saudi Arabia"(PDF). In Abu-Zinada, A. H.; Goriup, P. D.; Nader, I. A (eds.).Wildlife conservation and development in Saudi Arabia. 3. Riyadh: National Commission for Wildlife Conservation and Development Publishing. pp. 226–228.
  10. ^abcdHarrison, D. L. & Bates, P. J. J. (1991).The mammals of Arabia(PDF). Vol. 354. Sevenoaks, UK: Harrison Zoological Museum. pp. 167–170.
  11. ^Kitchener, A. C.; Breitenmoser-Würsten, C.; Eizirik, E.; Gentry, A.; Werdelin, L.; Wilting, A.; Yamaguchi, N.; Abramov, A. V.; Christiansen, P.; Driscoll, C.; Duckworth, J. W.; Johnson, W.; Luo, S.-J.; Meijaard, E.; O'Donoghue, P.; Sanderson, J.; Seymour, K.; Bruford, M.; Groves, C.; Hoffmann, M.; Nowell, K.; Timmons, Z. & Tobe, S. (2017)."A revised taxonomy of the Felidae: The final report of the Cat Classification Task Force of the IUCN Cat Specialist Group"(PDF).Cat News (Special Issue 11):73–75.
  12. ^Soultan, A.; Attum, O.; Hamada, A.; Hatab, E.-B.; Ahmed, S. E.; Eisa, A.; Sharif, I. A.; Nagy, A.; Shohdi, W. (2017). "Recent observation for leopardPanthera pardus in Egypt".Mammalia.81 (1):115–117.doi:10.1515/mammalia-2015-0089.S2CID 90676105.
  13. ^abcJudas, J.; Paillat, P.; Khoja, A.; Boug, A. (2006)."Status of the Arabian leopard in Saudi Arabia"(PDF).Cat News (Special Issue 1):11–19.
  14. ^Thesiger, W. (1949). "A Further Journey across the Empty Quarter".The Geographical Journal.113 (113):21–44.Bibcode:1949GeogJ.113...21T.doi:10.2307/1788902.JSTOR 1788902.
  15. ^"Arabian Tahr gets royal protection".WWF. 2009. Archived fromthe original on 2018-09-13. Retrieved2018-03-30.
  16. ^abcEdmonds, J.-A.; Budd, K. J.; Al Midfa, A. & Gross, C. (2006)."Status of the Arabian Leopard in United Arab Emirates"(PDF).Cat News (Special Issue 1):33–39.
  17. ^abSpalton, A.; Hikmani, H. D. a.; Willis, D. & Said, A. S. B. (2006)."Critically endangered Arabian leopardsPanthera pardus nimr persist in the Jabal Samhan Nature Reserve, Oman".Oryx.40 (3):287–294.doi:10.1017/S0030605306000743.
  18. ^Mazzolli, M. (2009). "Arabian LeopardPanthera pardus nimr status and habitat assessment in northwest Dhofar, Oman (Mammalia: Felidae)".Zoology in the Middle East.47:3–12.doi:10.1080/09397140.2009.10638341.S2CID 53510526.
  19. ^abAl Jumaily, M.; Mallon, D. P.; Nasher, A. K. & Thowabeh, N. (2006). "Status Report on Arabian Leopard in Yemen".Cat News. Special Issue 1:20–25.
  20. ^Nowell, K. & Jackson, P. (1996)."LeopardPanthera pardus".Wild cats: status survey and conservation action plan. Gland: IUCN/SSC Cat Specialist Group. Archived fromthe original on 2013-02-23. Retrieved2019-07-04.
  21. ^Perez, I.; Geffen, E. & Mokady, O. (2006)."Critically Endangered Arabian leopardsPanthera pardus nimr in Israel: estimating population parameters using molecular scatology".Oryx.40 (3):295–301.doi:10.1017/s0030605306000846.
  22. ^Granit, B. (2016)."Once there were Leopards". BirdLife Israel.
  23. ^Qarqaz, M. & Abu Baker, M. (2006). "The Leopard in Jordan".Cat News. Special Issue 1:9–10.
  24. ^Kingdon, J. (1990).Arabian Mammals. London: Academic Press Ltd.
  25. ^abSunquist, M. E.; Sunquist, F. (2002).Wild Cats of the World. Chicago: University of Chicago Press.ISBN 0-226-77999-8.
  26. ^Budd, K. (2001). "Arabian Leopards: New Hope is Born".Arabian Wildlife. 2000/2001:8–9.
  27. ^Breitenmoser, U. (2006).7th Conservation Workshop for the Fauna of Arabia 19–22 February: Workshop report. Sharjah, United Arab Emirates: Breeding Center for Endangered Arabian Wildlife.
  28. ^Mallon, D.P.; Budd, K. (2011)."Arabian LeopardPanthera pardus nimr (Hemprich & Ehrenberg, 1833)"(PDF).Regional Red List status of carnivores in the Arabian Peninsula. Gland and Sharjah, UAE: IUCN. pp. 13–15.
  29. ^Environment and Protected Areas Authority (2002).Conservation Assessment and Management Plan (CAMP) for the threatened fauna of Arabia's Mountain Habitat. Final Report. Sharjah, UAE: EPAA.
  30. ^Anonymous (2010)."Wildlife skins for sale after Haj – Saudi Arabia"(PDF).Wildlife Times.27:13–14.
  31. ^Zafar-ul Islam, M.; Boug, A.; Judas, J.; As-Shehri, A. (2018). "Conservation challenges for the Arabian Leopard (Panthera pardus nimr) in the Western Highlands of Arabia".Biodiversity.19 (3–4):188–197.Bibcode:2018Biodi..19..188Z.doi:10.1080/14888386.2018.1507008.S2CID 134163948.
  32. ^"The environmental impacts of the Israeli West Bank Barrier" by Michael L. Thomas and Karen E. Jenni.
  33. ^Sanborn, C.; Hoogstral, H. (1953). "Some mammals of Yemen and their parasites".Fieldiana Zoology.34: 229–.
  34. ^Rushby, K. (2011).Saving the Leopard (Documentary). Qatar: Al Jazeera.
  35. ^Anderson, D.; Grove, A. (1989).Conservation of Africa: People, Politics and Practice. New York:Cambridge University Press.
  36. ^Bailey, T. N. (1993).The African leopard: Ecology and Behavior of a Solitary Felid. New York:Columbia University Press.
  37. ^"Arabian leopard conservation project in Yemen".Rewilding Foundation. Retrieved2019-09-05.
  38. ^Shirka, H. (2019)."Al Ula conservation project can help Arabian leopards come roaring back".The National. Retrieved2019-09-05.
  39. ^Al-Khudair, D. (2019)."$20 million deal signed to save Arabian leopard population".Arab News. Retrieved2019-09-05.
  40. ^Atzeni, L.; Ilany, A.; Geffen, E.; Cushman, S. A.; Kaszta, Ż.; Macdonald, D. W. (2024)."Reviving the Arabian leopard: Harnessing historical data to map habitat and pave the way for reintroduction".Biological Conservation.291 110440.Bibcode:2024BCons.29110440A.doi:10.1016/j.biocon.2024.110440.
  41. ^Edmonds, J. A.; Budd, K. J.; Vercammen, P. & al Midfa, A. (2006). "History of the Arabian leopard Captive Breeding Programme".Cat News. Special Issue 1:40–43.
  42. ^Budd, J.; Leus, K. (2011)."The Arabian LeopardPanthera pardus nimr conservation breeding programme".Zoology in the Middle East.54 (Supplement 3):141–150.doi:10.1080/09397140.2011.10648905.
  43. ^"Sharjah's new wildlife centre opens to tourists". Sharjah Update. 2016.Archived from the original on 12 May 2019. Retrieved5 December 2018.
  44. ^De Haas van Dorsser, F. J.; Thowabeh, N. S.; Al Midfa, A. A.; Gross, C. (2001).Health status of zoo animals in Sana'a and Ta'izz, Republic of Yemen(PDF) (Report). Sana'a, Yemen:Breeding Centre for Endangered Arabian Wildlife, Sharjah; Environment Protection Authority. pp. 66–69. Archived from the original on May 5, 2019. Retrieved2019-05-05.
  45. ^"Successful birth of Arabian leopard cubs 'new beacon of hope' in Saudi bid to save species from extinction: Culture minister".Arab News. 2019. Retrieved2019-09-05.
  46. ^Read, Johanna (2022-02-09)."It's Arabian Leopard Day And AlUla, Saudi Arabia Is Working To Rewild The Endangered Species".Forbes. Retrieved2024-01-19.
  47. ^KOSSAIFY, EPHREM (2023-06-22)."UN recognition of Arabian Leopard Day a 'major triumph for KSA,' conservationist says".Arab News. Retrieved2024-01-19.
  48. ^"Arabian Leopard Day: Saudi Arabia's $25m fund will rewild critically-endangered species".The National News. 2023-02-09. Retrieved2024-01-19.
ExtantCarnivora species
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Panthera pardus nimr
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