| Amazon biome | |
|---|---|
 Anavilhanas National Park, Amazonas, Brazil | |
 Amazon biome (white outline)[1] and Amazon basin (light blue outline) | |
| Ecology | |
| Biome | Amazonia |
| Geography | |
| Area | 6,700,000 km2 (2,600,000 sq mi) |
| Countries | |
| Coordinates | 2°58′S64°36′W / 2.97°S 64.60°W /-2.97; -64.60 |
| Part ofa series on the |
| Amazon rainforest |
|---|
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Biodiversity |
TheAmazon biome (Portuguese:Bioma Amazônia) contains theAmazon rainforest, an area oftropical rainforest, and otherecoregions that cover most of theAmazon basin and some adjacent areas to the north and east. Thebiome containsblackwater andwhitewater flooded forest, lowland and montaneterra firma forest, bamboo and palm forest, savanna, sandy heath and alpine tundra. Some areas of the biome are threatened by deforestation for timber and to make way for pasture or soybean plantations.
The Amazon biome has an area of 6,700,000 square kilometres (2,600,000 sq mi).[2][a]The biome roughly corresponds to the Amazon basin, but excludes areas of theAndes to the west andcerrado (savannah) to the south, and includes lands to the northeast extending to the Atlantic ocean with similar vegetation to the Amazon basin.[2]J. J. Morrone (2006) defines the Amazonian subregion in this broader sense, divided into thebiogeographical provinces of Guyana, Humid Guyana, Napo, Imeri, Roraima, Amapá, Várzea, Ucayali, Madeira, Tapajós-Xingu, Pará, Yungas and Pantanal.[4][5]TheWorld Wildlife Fund takes a similar scope, where the Amazon biome includes theGuiana Shield rain forests in the north and theChiquitano dry forests of Bolivia.[6]
The biome covers parts of Brazil, Bolivia, Peru, Ecuador, Colombia, Venezuela, Guyana, Suriname and French Guiana.[2]In Brazil the biome covers more than 4,100,000 square kilometres (1,600,000 sq mi) and covers all or parts of the states ofAcre,Amazonas,Roraima,Rondônia,Pará,Amapá,Maranhão,Tocantins andMato Grosso.[7]The Amazon biome covers 49.29% of Brazil.[8]16% of the biome is in Peru.As of 2015 about 23.4% of Peru's Amazon biome was protected, but of this less than half was fully protected.[9]
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Much of the terrain of the Amazon biome, particularly around the rivers, is lowland plains.TheGuiana Shield is an area of highlands along the border between Brazil and Venezuela and Guyana.The southern Amazonian highlands cross parts of Rondonia and Mato Grosso and the southern parts of Amazonas and Para.[10]
The Amazon basin is crossed by ridges or "paleoarches" that connect theGuiana andCentral Brazil shields and divide it into geological sub-basins.They are the Iquitos or Jutai Arch in Peru andAcre, the Carauari Arch across theRio Negro andSolimões, the Purus Arch to the west ofManaus, the Monte Alegre Arch to the west of theTapajós and the Gurupa Arch to the west ofMarajó. Under the Paleoarch model, paleobasins between the arches form centers for biological diversification.Thus the Iquitos arch is considered the main reason for the different species of frogs and rodents and different forest types on either side of the ridge.[11]
The soil is generally very poor in nutrients, and areas that have been deforested are often unsuitable for agriculture or pasture.[12] A large part of thephosphorus necessary for life is blown by the wind from Africa; asdiatomeous dust from theBodélé Depression[13] and as smoke due to biomass burning in Africa.[14]There are wide regional variations in soil types.Thus 20% of theRio Negro basin is covered bypodzols and 55% byacrisols andferralsols, with the remainder covered by alluvial and litholic soils and scattered areas of hydromorphicplinthosols.In the biome as a whole podzols cover just 136,000 square kilometres (53,000 sq mi), or 2.7% of the area.[15]
In Brazil the average temperature of the biome is 22 to 26 °C (72 to 79 °F) and average rainfall is 2,300 millimetres (91 in), but there are wide variations from one region to another.[16]The biome as a whole has annual rainfall from 1,500 to 3,000 millimetres (59 to 118 in), about half of which is carried by winds from the Atlantic, and the other half fromevapotranspiration within the biome.There are wide variations in total rainfall and distribution of rainfall throughout the year.[2]
The Amazon watershed covers about 5,846,100 square kilometres (2,257,200 sq mi).[12]TheAmazon River accounts for 15–16% of the total water discharged by rivers into the oceans of the world.[2]Rivers may beblackwater,whitewater orclearwater.Thus theRio Negro ("Black River") has clear, jet-black water caused by decomposition of organic matter in swamps along its margins, combined with low levels of silt.TheRio Branco ("White River") and the Amazon itself have yellowish waters loaded with silt.[17]The Tahuayo River in theTamshiyacu Tahuayo Regional Conservation Area of Peru is classed as a blackwater river, but often has similar chemistry to the whitewater rivers of the region since it is in the Amazon River floodplain, and receives water from the Amazon.[18]
The Amazon and its major tributaries such as theXingu,Tapajós,Madeira,Purus andRio Negro form barriers to thegeodispersal of plants, animals and even insects. Thus thewhite-fronted capuchin (Cebus albifrons) andhairy saki (Pithecia hirsuta) are found west of the Tapajós, while thewhite-nosed saki (Chiropotes albinasus) is only found east of the river.[19]TheWorld Wildlife Fund divides the biome intoecoregions, often defined as the regions lying between major tributaries of the Amazon.
Most of the interior of the Amazon basin is covered by rainforest.[6]The dense tropical Amazon rainforest is the largest tropical rainforest in the world.[2]It covers between 5,500,000 and 6,200,000 square kilometres (2,100,000 and 2,400,000 sq mi) of the 6,700,000 to 6,900,000 square kilometres (2,600,000 to 2,700,000 sq mi) Amazon biome.The somewhat vague numbers are because the rainforest merges into similar biomes across its boundaries.[20]The rainforest isTropical Broadleaf Forest, so-called because most of the trees have broad leaves.[12]The basin also holds flooded riparian forest orvárzea, seasonal forest andsavanna. Seasonal forest covers much of the southeast border, with marked dry seasons when there are frequent fires.[6]The Amazon biome contains areas of other types of vegetation including grasslands, swamps, bamboos, and palm forests.[2]
There are 53 majorecosystems and more than 600 types of land and freshwaterhabitat.Of the ecosystems, 34 are forest areas covering 78% of the biome, 6 are Andean covering 1.5%, 5 are floodplains covering 5.83%, 5 are savanna covering 12.75% and two are tropical steppes covering 1.89%.The Brazilian Amazon holds 30 of the 53 ecosystems, of which 19 are forests with 77.5% of the area.[16]The borders of the biome holdecotones where it blends into other biomes such as the cerrado.[7]
Within and across the ecosystems of the biome there is huge biological diversity.One source says there are an estimated 60,000 plant species, of which 30,000 are endemic.[3]Another says there are 30,000 to 50,000 plant species.[16]The Brazilian Ministry of the Environment as of January 2013 listed 2,500 species of trees and 30,000 species of plants.[10]There are 1,400 species of fish, 163 amphibians, 387 reptiles and more than 500 mammals including 90 primates.87% of the amphibians, 62% of reptiles, 20% of birds and 25% of mammals are endemic to the biome.[16]109 species of lizards andamphisbaena reptiles are known to be present, and 138 species of snakes.Of the 1,300 bird species 20% are endemic and 8.4% are endangered.[21]
A relatively small area may contain a variety of ecoregions.Thus thePico da Neblina National Park in the north of Amazonas, Brazil containscampinarana, dense rainforest and contact between campinarana and rainforest.[22]The vegetation formations include terra firme forests andigapós.[22]Submontane forests climb the first steps of the Guiana Plateau to about 1,000 metres (3,300 ft), followed by montane forests.Lichens and bromeliads are found up to 2,000 metres (6,600 ft).There is alsoalpine tundra in the tabular plateaus.[22]
The Amazon basin contains several large areas of moist forest, collectively called the Amazon rainforest.These are theCaquetá,Japurá–Solimões–Negro,Juruá–Purus,Madeira–Tapajós,Napo,Purus–Madeira,Solimões–Japurá,Southwest Amazon,Tapajós–Xingu,Tocantins–Araguaia–Maranhão,Uatuma–Trombetas,Ucayali, andXingu–Tocantins–Araguaia moist forests.Each has distinctive vegetation.[23]In the broad definition favored by theWorld Wide Fund for Nature (WWF) the Amazon biome would also include theTepuis,table-top mountains with unique vegetation, theNegro-Branco,Guianan Highlands,Guianan piedmont and lowland andGuianan moist forests and theGuianan savanna.[24][2]
In the southwest Amazon at least 161,500 square kilometres (62,400 sq mi) of forest are dominated bybamboos.These occur in areas where there has been recent tectonic uplift combined with fast mechanical erosion and poor drainage.In each patch of bamboos all the plants flower every 27–28 years, producing huge numbers of seeds, before dying.[25]TheChandless State Park is in the centre of the bamboo forest region of the south western Amazon biome, and has three endemic species of theGuadua genus of bamboo. Vegetation is classified as forest with palms, forest with bamboo, alluvial forest with bamboo and dense forest.[26]
BeforeGhillean Prance's 1979 study, the local people and many scientists called the permanently waterlogged swamp on the lower Amazon "igapó" and all types of periodically flooded land "várzea".[27]A more recent definition, from Prance, is:[28]
| Type of flooding | Type of water | Name | Alternative names |
|---|---|---|---|
| Seasonal | Whitewater | Seasonal várzea | |
| Blackwater | Seasonal igapó | ||
| Tidal | Salt water | Mangrove | |
| Fresh water back-up | Tidal várzea | ||
| Irregular rainfall | Floodplain forest | Riparian forest along low order rivers | |
| Permanent | Whitewater | Permanent swamp forest | várzea swamp forest, várzea chavascal |
| Blackwater and clearwater | Permanent igapó | Long-term inundated forests |
Major flooded areas include theIquitos,Purus,Monte Alegre,Gurupa andMarajó várzeas.The Majaró várzea is at the mouth of the Amazon and is affected by both freshwater and tidal flows.[23]
Campina is open forest on sandy soil where sunlight can reach the ground.More than half the species of orchid in the Amazon lowlands are found in this type of forest.[29]The terms campina andcampinarana both describe white sand savannas that are very poor in nutrients.They may be flooded periodically or seasonally, in which case the roots suffer from lack of aeration.[30]The vegetation is stunted.[31]For some,campina is distinguished as being completely treeless.[32]The term "campinarana" is used for the transition betweencampina and the rainforest.[33]
Campinarana is found in isolated areas of theRio Negro andRio Branco basins in the north of Brazil.[34]Patches of campinarana, which may cover several thousand square kilometres, are found in the transitional region from the Guyana Shield to the Amazon basin.[35]Campinarana is typically found on leached white sands around circular swampy depressions in lowland tropical moist forest.The soil is low in nutrients, with highly acidic humus.[34]Campinarana includes savannah, scrub and forests.[34]The savannah is mainly composed of grasses and lichens, found in the wet plains beside lakes and rivers.[36]The scrub has bare sand, herbaceous plants, shrubs and trees less than 7 metres (23 ft) high.[34]
The nutrient-poor Japurá-Solimões-Negro moist forest is similar to the Rio Negro Campinarana.[23]
TheNortheastern Brazil restingas are coastal dune habitats that extend along the coast of northeastern Brazil, interspersed with lagoons, mangroves and patches ofcaatinga savanna.The land behind the dunes may include dwarf palms, bromeliads, ferns, shrubs, grasses and scrub trees.The more exposed areas mainly hold medium-tall grasses and scrub trees, while sheltered areas hold patches of cactus and low dry thicket.[37]Fauna includemarmosets andjaguarundis,proboscis bats,lesser sac-winged bats,bulldog bats, andDavy's naked-backed bats,wood stork,roseate spoonbill,white-necked heron,great egret,cattle egret,black-crowned night heron, andNeotropic cormorant.The ecoregion has various endemic species.TheLençóis Maranhenses National Park protects a large area of this habitat.[37]
Of the 1,149 federal and state protected areas in Brazil in 2014, 247 covering 1,100,000 square kilometres (420,000 sq mi) were in the Amazon biome and accounted for 73% of the total area protected at the federal and state levels in Brazil.Federal units covered 587,000 square kilometres (227,000 sq mi) and state units covered 523,000 square kilometres (202,000 sq mi).[38]These included fully protectednational andstate parks (22%),ecological stations (9%),biological reserves (4%) andwildlife refuges as well as sustainable usenational andstate forests (26%),environmental protection areas (17%),extractive reserves (12%),sustainable development reserves (10%), andareas of ecological interest.[39]More than 15,000 square kilometres (5,800 sq mi) of the Brazilian Amazon biome was deforested in 2008–12, of which only 6% took place within protected areas.[40]71% of federal and state protected areas in Brazil suffered no deforestation in this period.[41]
Although theWorld Economic Forum ranks Brazil first in the world in terms of nature tourism potential, the country is 52nd in tourism competitiveness when factors such as infrastructure are considered.Low public use, in part due to the inaccessibility of the Amazon national and federal parks, mean that they fail to support the local economies and therefore lack support from society, especially the local communities.[42]The national and state forests have low levels of formal sustainable logging contracts, theTapajós National Forest in Pará being an exception.[43]Brazil has strong systems for monitoring land cover and deforestation, but as of 2014 64% of managers of protected areas said they had not monitored biodiversity in the previous five years, and more than half said they did not have tools to monitor social and environmental development.[44]
The "Arc of Deforestation" is in the eastern and southern borders of the biome.[45]As of 2006 about 16% of the Amazon biome in Brazil had been deforested.[46]Satellite images show that in the 2006–11 period total deforestation in the Amazon biome was 45,100 square kilometres (17,400 sq mi), of which 34,700 hectares (86,000 acres) were in the three soy-producing states of Mato Grosso, Para and Rondonia.A 2009 survey showed that of 620 recently deforested areas 203, or 32%, had been converted to pasture while 12, or less than 1%, were being used for soybeans.[47]
The biome is not the same as theAmazônia Legal, which covers a larger area of Brazil.[48]The Amazon biome covers about 82% of the Legal Amazon if cerrado and campinarana are excluded.[49]Defenders of soybean farming point out that the soybean farms in the Amazonia Legal mostly occupy cerrado areas, not the Amazon biome.[48]A statement byCargill in 2006 said, "Soy occupies less than 0.6 percent of the land in the Amazon biome today, and most of that soy is grown on the fringes of the Amazon biome in the transitional area between theCerrado and the forest.[50]A 2010 assessment of soybean farmers associated with theGrupo André Maggi found that 62.55% of the properties produced 78.09% of the crop in the cerrado biome, while 37.45% produced 21.91% of the crop in the Amazon biome.[51]
In 2000 the Brazilian government banned creation of sugarcane plantations in the Amazon biome.[52]In 2010 a presidential decree made low-interest credit available to oil palm development, but only on land that had been deforested before 2008.This may prove an environmentally sustainable economic solution.[53]
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